Echinocereus morricalii – Echinocereus reichenbachianus

Echinocereus morricalii Riha 1975:75.
TL: Mexico, Nuevo Leon, Barranca de las Garrapatassouth of
Monterrey, D. B. Morrical (PRC 322).
Synonyms:
Echinocereus viereckii var. morricalii (Riha) Taylor 1985b:93.
Echinocereus blanckii var. inermis* Heyer 1989:26 pro syn.
Echinocereus viereckii ssp. morricalii (Riha) Taylor 1997:10.
*Echinocereus multiangularis = Haageocereus sp.
*Echinocereus mutitangularis var. limensis = Haageocereus sp.
*Echinocereus multiangularis var. maletianus = ?
*Echinocereus multiangularis var. pallidior  = ?
Echinocereus multicostatus Cels ex Rümpler 1885:834.
TL: unknown
Synonyms:
Echinocereus leeanus var. multicostatus (Cels ex Rümpler)
Schumann 1897:289.
Echinocereus triglochidiatus var. multicostatus (Cels ex
Rümpler) Marshall 1953:67.
Note: this plant is often considered a synonym of
          Echinocereus leeanus, but its may be of South American
origin and not an Echinocereus.
Echinocereus munzii (Parish) Benson 1941:361.
= Cereus munzii Parish 1926:48.
TL: California, Riverside Co., 2 miles below Kenworthy, 1400m,
Munz & Johnson 5570, 21 May 1922(UC).
Synonyms:
Echinocereus engelmannii var. munzii (Parish) Pierce & Fosberg 1933:123.
Echinocereus neomexicanus Standley 1908:87-88.
TL: New Mexico, Doña Ana Co., mesa west of Organ Mtns.,
Standley 383, 1907(MO).
Synonyms:
Echinocereus triglochidiatus var. neomexicanus (Standley)
Standley ex Marshall & Bock 1941:118.
Cereus neomexicanus (Standley) Tidestrom 1941:298
Echinocereus polyacanthus var. neomexicanus* Weniger1970:44 nom. inval.
Echinocereus coccineus var. neomexicanus* Parker 1996:4 nom. nud.
Note:This plant has been confused with Echinocereus rosei
          under the name Echinocereus triglochidiatus var.  neomexicanus [Benson] and
         Echinocereus polyacanthus  [Bravo 1991:24].It is probably a natural hybrid of E. 
          rosei and E. chloranthus.
Echinocereus neocapillus (Weniger) Blum & Lange in Blum et al. 1998:219.
  = Echinocereus chloranthus var. neocapillus.
Echinocereus neoroemeri* (Mühlenpfordt) Ito 1971:227. nom. illegit. 
  = Echinocereus roemeri (Mühlenpfordt) Rydberg
Wilcoxia nerispina* Backeberg 1960:2078 nom. invalid. 
  = Echinocereus schmollii.
Echinocereus nicholii (Benson) Parfitt 1987:157-158
  = Echinocereus engelmannii var. nicholii.
Echinocereus nicholii fa. guaymasensis* Parker 1996:1 nom. nud.
Echinocereus nicholii ssp. llanuraensis Rutow 1995:61-71.
TL: Mexico, Sonora, Guaymas, 300m (ZSS # A 16936).
Echinocereus nicholii fa. llanuraensis* Parker 1996:1 nom. nud. 
  = Echinocereus nicholii ssp. llanuraensis
Echinocereus nigrispinus* Köhres Catalog 1979. nom. nud.
Echinocereus nivilus* Heyer 1989:26  nom. nud. [= Lau 1101].
Echinocereus nivosus Glass & Foster 1978a:18-19.
TL: Mexico, Coahuila, Southwest of General Cepeda, pass 3 km.
north of El Cinco, Glass & Foster 3764, Feb. 1972 (POM).
Note: Echinocereus albatus* Backeberg 1960:2007 nom. inval.
(no type designated) is usually considered a synonym of this
species.
Echinocereus nivosus fa. albatus* Parker 1996:3 nom. nud.
= Echinocereus albatus*.
Echinocereus noctiflorus* Hort. ex Backeberg 1960:1991 pro syn. 
  = Echinocereus cucumis.
Echinocereus nocturniflorus* Hort. ex Backeberg 1960:1991 pro syn. 
  = Echinocereus cucumis.
Echinocereus nogalensis* Haage 1900:159 nom. nud.
  [Perhaps = Echinocereus pectinatus var. rigidissimus].
Echinocereus ochoterenae Ortega 1928:88-90.
TL: Mexico, Sinaloa, near Concordia, Cerro de la Cobriza, J.G.
Ortega, 24 May 1927 (MEXU?).
Synonyms:
Echinocereus subinermis var. ochoterenae (Ortega) Unger 1984:164.
Echinocereus subinermis ssp. ochoterenae (Ortega) Taylor 1997:10.
Note: Ortega described varieties „A“ and „B“, but did  not give them names.
Echinocereus octacanthus (Mühlenpfordt) Britton & Rose 1922:13.
= Echinopsis octacantha Mühlenpfordt 1848:19.
TL: Northern Texas [San Saba River according to Engelmann],
Roemer.
Synonyms:
Cereus octacanthus (Mühlenpfordt)Coulter 1896:395.
Echinocereus coccineus var. octacanthus* Boissevain & Davidson 1940:39 nom. illegit.
Echinocereus triglochidiatus var. octacanthus* Mühlenpfordt ex Marshall & Bock
1941:118. nom. illegit.
Note: The proper application of this name is uncertain. If it can be shown to be one of
the members of the  Echinocereus coccineus group it would have priority
since it was published first. This would cause considerable nomenclatural changes.
Echinocereus oklahomensis Lahman 1950:128.
[Originally published in Lahman 1935:141 without Latin diagnosis].
TL: Oklahoma, Wichita Mtns., Lahman (MO).
Note: Usually consisdered a synonym of Echinocereus baileyi.
Echinocereus orcuttii Rose ex Orcutt 1926:4.
TL: Mexico, Baja California, Cape Colset, Orcutt (?).
Note: Usually consisdered a synonym of Echinocereus maritimus.
Echinocereus ortegae Rose ex Ortega 1929:24.
TL: Mexico, Sinaloa, Cuenca del Rio Tamazula, Sianori, 800m., Ortega (?).
Note: Taylor (1993) lists Echinocereusscheeri var. koehresianus as a synonym
of this species.
Echinocereus ortegae ssp. koehresianus (Frank) Rischer & Frank 1996:93
    = Echinocereus scheeri var. koehresianus.
Echinocereus pacificus* Haage 1892:118 nom.nud.
Echinocereus pacificus (Engelmann ex Orcutt) Britton & Rose 1922:12.
= Cereus phoeniceus var. pacificus Engelmann ex Orcutt 1886:46.
TL: Mexico, Baja California Norte, Rio San Carlos, Parry et al., 25 Jan. 1883 (MO).
Synonyms:
Cereus pacificus (Engelmann ex Orcutt) Coulter 1896:397.
Echinocereus triglochidiatus var. pacificus (Engelmann ex Orcutt) Engelmann ex 
Marshall & Bock 1941:118.
Echinocereus polyacanthus var. pacificus (Engelmann ex Orcutt) Taylor 1984:160.
Echinocereus polyacanthus ssp. pacificus (Engelmann ex Orcutt) Breckwoldt 1996:15.
Echinocereus pacificus ssp. mombergerianusBlum, Rischer & Rutow in Blum et al. 1998:418. = Echinocereus mombergerianus.
Echinocereus palmeri Britton & Rose 1922:34.
TL: Mexico, Chihuahua, hill near Cd. Chihuahua, E. Palmer 121,
Apr. 1908 (US).
Echinocereus palmeri
ssp. escobedensis Rischer (KuaS 54 (5) 2003)
TL:
Durango, MX. Lau 1565, 18.3.1988 ex cult. coll. Rischer
Echinocereus pamanesiorum Lau 1981:36-41.
TL: Mexico, Zacatecas, Rio Huaynamota, bridge between
Huejuquilla & San Juan Capistrano, Lau 1247 (MEXU).
Echinocereus pamanesiorum ssp. bonatzii (Römer) Römer 1997:45.
= Echinocereus bonatzii.
Echinocereus papillosus Linke ex Rümpler 1885:783.
TL: unknown
Synonyms:
Cereus papillosus (Linke ex Rümpler) Berger 1905:80.
Echinocereus blanckii var. papillosus (Linke ex Rümpler) Benson 1969d:126.
Echinocereus berlandieri var. papillosus (Linke ex Rümpler) Benson 1976:59.
Note: Benson uses the names Echinocereus blanckii var. angusticeps
and Echinocereus berlandieri var. angusticeps for this plant. Usually listed as
Echinocereus papillosus Linke ex Rümpler 1885:783.
Echinocereus papillosus var. angusticeps (Clover) Marshall & Bock 1941:119
= Echinocereus angusticeps.
Echinocereus papillosus ssp. angusticeps* Blum 1997 Seed  List nom. nud.
  = Echinocereus angusticeps.
Echinocereus papillosus var. giganteus* Bailey 1941:270 nom. nud.
Echinocereus papillosus var. rubescens Hildmann ex Schelle
1926:165.
TL: unknown
Echinocereus parkeri Taylor 1988:73.
TL: Mexico, Nuevo Leon, between La Poza & Puerto de los Pastores,
Hansen et al. 3863 (MEXU).
Synonyms:
Echinocereus stramineus var. parkeri* Seed list 1996 –  British Cactus and Succulent 
        Journal.
Echinocereus parkeri ssp. arteagensis Blum & Lange in Blum et al. 1998:175.
TL: Mexico, Coahuila – Nuevo Leon Border, Arteaga Canyon, 2250 m,
        U. Guzman in Sánchez- Mejorada 3807 (MEXU).
Echinocereus parkeri fa. barcena* Parker 1996:3 nom. nud. 
  = Echinocereus barcena.
Echinocereus parkeri var. gonzalezii Taylor 1988:74.
TL: Mexico, Tamaulipas, Sierra de las Vacas, 4km NE of San Jose
d’Llano, 1750m, Gonzalez Medrano 8515 (MEXU).
Echinocereus parkeri ssp. gonzalezii BTaylor) Taylor 1997:9.
   = Echinocereus parkeri var. gonzalezii.
Echinocereus parkeri var. mazapilensis* K. Neumann 1993 nom. prov.
Kaktusblüte p. 55.
Echinocereus parkeri ssp. mazapilensis Blum & Lange in Blum et al. 1998:172.
TL: Mexico, Zacatecas, 0.3 km from El Cobre near Mazapil, 2750 m,
Sánchez – Mejoradaet al. 4041 (MEXU).
Echinocereus paucispinus (Engelmann) Loder 1882:445.
= Cereus paucispinus Engelmann 1856:285.
[fuller description Engelmann 1859:34].
TL: Texas, San Pedro River, Wright, 1849 (MO).
Synonyms:
Echinocereus paucispinus* Engelmann ex Rümpler 1885:794
nom. illegit.
  Echinocereus triglochidiatus var. paucispinus (Engelmann) Engelmann ex
          Marshall & Bock 1941:117.
Echinocereus coccineus var. paucispinus (Engelmann) Ferguson 1989:222.
Echinocereus coccineus ssp. paucispinus Blum, Lange & Rutow
in Blum et al. 1998:433.
           Note:Usually cited as Echinocereus paucispinus Engelmann ex
Rümpler 1885:794
Echinocereus paucispinus var. flavispinus* Haage 1892:118 nom. nud.
Echinocereus paucispinus var. flavispinus* Walton 1899b:18 nom. nud.
Echinocereus paucispinus var. flavispinus Hort. ex Schelle 1926:178.
TL: unknown
Echinocereus paucispinus var. fulvispinus* Haage 1897:12 nom. nud.
Echinocereus paucispinus var. fulvispinus Bowles 1908:28.
TL: unknown
Echinocereus paucispinus var. gonacanthus (Engelmann & Bigelow) Schumann 1897:281
    = Echinocereus gonacanthus.
Echinocereus paucispinus var. hexaedrus (Engelmann & Bigelow) Schumann 1897:281
    = Echinocereus hexaedrus.
Echinocereus paucispinus var. nigrispinus* Haage 1897:12 nom. nud.
Echinocereus paucispinus var. nigrispinus Bowles 1908:28.
TL: unknown
Echinocereus paucispinus var. triglochidiatus* Schumann 1897:281  nom. illegit. 
   = Echinocereus triglochidiatus.
Echinocereus pectinatus (Scheidweiler) Engelmann 1848a:110.
= Echinocactus pectinatus Scheidweiler 1838:492.
TL: Mexico, San Luis Potosi, Villa del Peñasco, Galeotti (?).
Synonyms:
Echinopsis pectinata (Scheidweiler) Fennel 1843:282.
Cereus pectinatus (Scheidweiler) Engelmann 1849:50.
Note:Echinocereus cernosus* Fric 1929:24 nom. nud. is
listed by Backeberg as a synonym of this species.
Echinocactus pectiniferusLemaire 1839:25 is usually
considered a synonym of this species.
Echinocereus pectinatus var. adustus* Haage 1859:19 nom. nud. 
Echinocereus pectinatus var. adustus [mispelled adjustus](Engelmann) Daul 1890:78
= Echinocereus adustus.
Note: Usually ascribed to Schumann 1897:271.
Echinocereus pectinatus fa. adustus (Engelmann) Voss 1894:378 = Echinocereus adustus.
Echinocereus pectinatus var. adustus fa. castaneus (Engelmann) Engelm. ex Schelle
1926:174   = Echinocereus caespitosus var. castaneus.
Echinocereus pectinatus var. armatus* Engelmann ex Seitz 1870:11 nom. nud.
Echinocereus pectinatus var. armatus (Poselger) Poselger ex 
  Rümpler 1885:818.
= Cereus pectinatus var. armatus Poselger 1853:134.
TL: Mexico, Nuevo Leon, near Monterrey. Poselger (?)
Synonyms:
Cereuspectinatus var. spinosus* Coulter 1896:357.
Echinocereus spinosus  Coulter ex Schelle 1926:174 pro syn.
Echinocereus armatus (Poselger) Poselger ex Backeberg & Knuth 1935:318.
Echinocereus caespitosus var. armatus (Poselger) Poselger ex Borg 1937:174.
Echinocereus reichenbachii var. armatus (Poselger) Taylor 1985b:133.
Echinocereus fitchii var. armatus* Mesa Garden Seed Catalog 1996 nom. nud.
Echinocereus reichenbachii ssp. armatus (Poselger) Taylor 1997:9.
Echinocereus fitchii ssp.armatus (Poselger) Blum et al. 1998:314
Echinocereus pectinatus var. bauerii Rother 1923:127.
TL: [no type locality or type mentioned]
Echinocereus pectinatus var. bristolii (Marshall) Marshall1956b:81-82
= Echinocereus bristolii.
Echinocereus pectinatus var. caespitosus (Engelmann) Schumann 1897:272
= Echinocereus caespitosus.
Echinocereus pectinatus var. candicans* Hort. ex Schelle 1907:132 nom. nud.
Echinocereus pectinatus var. candidissimus* Seitz 1878:11 nom. nud.
Echinocereus pectinatus var. castaneus* Seitz 1878:11 nom. nud.
Echinocereus pectinatus var. castaneus (Engelmann) Daul 1890:78
= Echinocereus caespitosus var. castaneus.
Echinocereus pectinatus fa. castaneus (Engelmann) Krainz 1967:50
= Echinocereus caespitosus var. castaneus.
*Echinocereus pectinatus var. centralis Coulter ex Schumann1897:271
error in determination.  = Echinomastus centrilis.
Echinocereus pectinatus var. chloranthus* Haage 1892:118 nom. nud.
Echinocereus pectinatus var. chrysacanthus Schumann 1897:272.
TL: Mexico.
Synonyms:
Echinocereus caespitosus var. chrysacanthus (Schumann)
Schumann ex Borg 1937:174.
Echinocereus pectinatus var. crassispinus Rünge ex Mathsson
1890:465.
TL: Mexico, Coahuila, Saltillo, Mathsson (?).
Echinocereus pectinatus var. ctenoides* Hort. ex Schelle1907:130 pro syn.
    = Echinocereus ctenoides.
Echinocereus pectinatus var. ctenoides* Weniger 1971:31 nom. inval. (no basionym given)
   = Echinocereus ctenoides.
Echinocereus pectinatus var. ctenoides (Engelmann) Weniger ex Frank 1991:135
= Echinocereus ctenoides.
Echinocereus pectinatus ssp. ctenoides (Engelmann) Frank 1997:136
= Echinocereus ctenoides.
Echinocereus pectinatus var. dasyacanthus (Engelmann) Engelmann ex Haage 1892:118
= Echinocereus dasyacanthus.
Echinocereus pectinatus var. dasyacanthus* Earle 1971:80 nom. illegit.  (later homonym)
   = Echinocereus dasyacanthus.
Echinocereus pectinatus var. dasyacanthus* Taylor 1984:179  nom. illegit. (later homonym)
   = Echinocereus dasyacanthus.
Echinocereus pectinatus var. grandicostatus* Schmoll ex Backeberg 1960:2043 nom. nud.
Echinocereus pectinatus var. grandis (Britton & Rose) Marshall 1955:68
= Echinocereus grandis.
Echinocereus pectinatus var. jamauvei *Fric ex Orcutt 1926:4 pro syn. 
  = Echinocereus castaneus.
Echinocereus pectinatus var. labouretianus*  Seitz 1870:11 nom. nud.
Echinocereus pectinatus var. laevior* (Monatsschrift für Kakteenkunde
  index 1912:56) nom. nud.
Echinocereus pectinatus var. minor (Engelmann) Benson 1968:125
for Echinocereus dasyacanthus var. minor*.
    = Echinocereus roetteri.
Echinocereus pectinatus var. neomexicanus (Coulter) Benson 1944:256
= Echinocereus dasyacanthus var. neomexicanus*.
Note: Benson uses this name for Echinocereus dasyacanthus.
Echinocereus pectinatus var. ordustus* Lakerveld Catalog. nom. nud.
Echinocereus pectinatus var. pailanus* Schwarz ex Backeberg 1960:2043 nom. nud.
Echinocereus pectinatus var. pailensis* Bailey 1941:271 nom. nud.
Echinocereus pectinatus var. pictus* Seitz 1870:11 nom. nud.
Echinocereus pectinatus var. rectispinus* Zimmerman1993:277 nom. nud. 
  = Echinocereus dasyacanthus var. rectispinus.
Echinocereus pectinatus var. reichenbachii (Terscheck ex Walpers) Werdermann 1930:t5
= Echinocereus reichenbachii.
Echinocereus pectinatus var. rigidissimus (Engelmann) Engelmann
  ex Rümpler 1885:818.
= Cereus pectinatus var. rigidissimus Engelmann 1856:279.
TL: Arizona, Sierra Pajarito, Schott 1855 (MO).
Synonyms:
Echinocereus pectinatus var. robustior* Hort. ex Hirscht
1897:95 pro syn.
Echinocereus candicans* Runge ex Haage 1900:157 pro syn.
Echinocereus rigidissimus (Engelmann) Hort. ex F. A. Haage 1897, „special
offer“catalog p. 13.
Echinocereus robustior* Britton & Rose 1922:28  pro syn.
Echinocereus dasyacanthus var. rigidissimus (Engelmann) Marshall & Bock 1941:119.
Echinocereus pectinatus fa. rigidissimus (Engelmann) Krainz 1967:50.
Note: Usually listed as Echinocereus rigidissimus Rose 1909a:293.
                Echinocereus nogalensis* Haage 1900:159 nom. nud.  is usually considered a
synonym of this plant.
Echinocereus pectinatus fa. rigidissimus (Engelmann) Krainz 1967:50.
=Echinocereus pectinatus var. rigidissimus.
Echinocereus pectinatus var. robustior* Hort. ex Hirscht 1897:95  pro syn.
  = Echinocereus pectinatus var. rigidissimus.
Echinocereus pectinatus var. robustus Bauer ex Wittmack 1890:513.
TL: Mexico, Sonora, Nogales.
Note: usually considered a synonym of Echinocereuspectinatus var. rigidissimus.
Echinocereus pectinatus var. rotatus (Linke) Linke ex Haage 1892:118
= Echinocereus rotatus.
Echinocereus pectinatus var. rubescens* Weniger 1970:21-22 nom. nud.
Echinocereus pectinatus var. rubispinus Frank & Lau in Frank
1982:32-35.
TL: Mexico, Chihuahua, Cañon de Barbarocas, near Campito in Sierra
Obscura, 1600-1800m, Lau 088 (ZSS AA 18-30).
Synonyms:
Echinocereus rigidissimus var. rubispinus (Frank & Lau) Taylor 1984:175.
Echinocereus rigidissimus ssp. rubispinus Taylor 1997:9.
Echinocereus pectinatus var. rungei* Hort. ex Schelle 1926:174  nom. nud.
Echinocereus pectinatus var. rufispinus* Daul 1890:78 nom. nud.
Echinocereus pectinatus var. rufispinus Schumann 1897:272.
TL: Mexico.
Synonyms:
Echinocereus caespitosus var. rufispinus (Schumann) Schumann ex Borg 1951:221.
Cereus pectinatus fa. rufispinus (Engelmann) Voss 1894:378
= Echinocereus rufispinus.
Echinocereus pectinatus var. scopulorum (Britton & Rose) Marshall 1955:61
= Echinocereus scopulorum.
Echinocereus pectinatus var. sonorensis* (Kaktusy 1973 according toEvanich 1989:111).
Cereus pectinatus var. spinosus* Coulter 1896:357 nom.illegit.
[an unnecessary new name for Echinocereus pectinatus var. armatus].
Echinocereus pectinatus var. steereae* Rümpler ex Weniger 1970:32  pro syn.
   = Echinocereus steereae.
Echinocereus pectinatus var. tamaulipensis Fric ex Schelle 1926:174.
TL: unknown
synonym:
Echinocereus caespitosus var. tamaulipensis (Fric) Fric ex Borg 1937:174.
Echinocereus pectinatus var. texensis* Horner ex Seitz 1870:11 nom. nud.
Echinocereus pectinatus var. texensis Hort. ex Rümpler 1885:817.
TL: Texas
Echinocereus pectinatus var. texensis* Hooker ex Schelle 1907:132  nom. nud.
Echinocereus pectinatus var. wenigeri Benson 1968:124.
TL: Texas, Valverde Co, near Langtry, Benson 16,521 (POM 311,338).
Echinocereus pectinatus ssp. wenigeri (Benson) Blum & Rutow in Blum et
al. 1998: 111=Echinocereus pectinatus var. wenigeri.
Echinocactus pectiniferus Lemaire 1839:25.
TL: unknown
Synonyms:
Cereus pectiniferus (Lemaire) Labouret 1853:320
Note: usually considered a synonym of Echinocereus pectinatus.
Echinocactus pectiniferus var. laevior Monville ex Lemaire 1839:26.
TL: unknown
Synonyms:
Echinopsis pectinata var. laevior (Monville ex Lemaire) Monville ex Förster 1846:365.
*Echinocereus pencilliformis Linke 1858
[of Bolivian origin and therefor not an Echinocereus].
Echinocereus pensilis (K. Brandegge) J. A. Purpus 1908:5.
= Cereus pensilis K. Brandegee 1904:192.
TL: Mexico, Baja California Sur, Sierra de Laguna, T.S. 
    Brandegee 246, Jan/Feb. 1890 (UC).
Synonyms:
Morangaya pensilis (K. Brandegee) Rowley 1974b:44-45.
Echinocereus pentalophus* Engelmann ex Haage 1859:20 nom. nud.
Echinocereus pentalophus* Lemaire 1868:56. nom. nud.
[Probably = Echinocereus pentalophus De Candolle ex Rümpler].
Echinocereus pentalophus (De Candolle) Lemaire ex Rümpler 1885:774.
= Cereus pentalophus De Candolle 1828:117.
TL: Mexico, Hidalgo, near Zimapan, Coulter (?).
Synonyms:
Cereus propinquus* Hort. Genev. ex Walpers 1843:275 pro syn.
Echinocereus pentalophus var. aureovariegatus* Ito 1952:134 nom. nud.
Echinocereus pentalophus var. ehrenbergii (Pfeiffer) Backeberg1960:2003
= Echinocereus ehrenbergii.
Echinocereus pentalophus var. leonensis (Mathsson) Taylor 1985b:78
= Echinocereus leonensis.
Echinocereus pentalophus ssp. leonensis (Mathsson) Taylor 1997:9
= Echinocereus leonensis.
Echinocereus pentalophus
var. leptacanthus* Haage 1859:20 nom. nud.
Echinocereus pentalophus var. leptacanthus* Lemaire ex Rümpler 1885:775 nom. illegit. 
    = Echinocereus leptacanthus.
Echinocereus pentalophus var. procumbens (Engelmann) Fournier 1935:25
= Echinocereus procumbens.
Echinocereus pentalophus fa. procumbens* Parker 1996:4 nom. nud. 
  = Echinocereus procumbens.
Echinocereus pentalophus ssp. procumbens (Engelmann) Blum & Lange
in Blum et al. 1998:129. = Echinocereus procumbens.
Cereus pentalophus var. radicans DeCandolle1828:117.
TL: Mexico, Hidalgo, near Zimapan, Coulter (?).
Synonyms:
Cereus propinquus* Hort. Genev. ex Walpers 1843:275 pro syn.
Echinocereus pentalophus var. radicans* Weniger 1970:51  nom. inval.
Echinocereus pentalophus var. radicans* Weniger 1970:51 nom. inval. 
(no basionym given). = Cereus pentalophus var. radicans.
Cereus pentalophus var. simplex De Candolle 1828:117.
TL: Mexico, Hidalgo, near Zimapan, Coulter (?).
Synonyms:
Cereus propinquus* Salm Dyck ex Otto:1833:366. nom. nud.
Cereus propinquus* De Candolle ex Walpers 1843:333 pro syn.
Echinocereus propinquus De Candolle ex Haage 1892:118.
Echinocereus pentalophus var. simplex* Weniger 1970:51nom. inval.
Echinocereus pentalophus var. simplex* Weniger 1970:51 nom. inval. (no basionym given).
= Cereus pentalophus
var. simplex.
Cereus pentalophus var. subarticulatus De Candolle 1828:117.
TL: Hidalgo, near Zimapan, Coulter (?).
Synonyms:
Cereus propinquus var. subarticulatus* Pfeiffer ex Förster 1846:373 pro syn.
Cereus leptacanthus De Candolle ex Salm-Dyck 1845:27.
Echinocereus leptacanthus Gaillard 1864:209.
Echinocereus pentalophus var. subarticulatus* Weniger 1970:51 nom. inval.
Echinocereus pentalophus var. subarticulatus* Weniger 1970:51 nom. inval.
    (no basionym given).  = Cereus pentalophus var. subarticulatus.
*Echinocereus pentlandii  = Echinopsis pentlandii.
Echinocereus perbellus Britton & Rose 1922:24-25.
TL: Texas, Big Springs, Rose & Standley 12215, 23 Feb.1910 (US 635015, 635016).
Synonyms:
Echinocereus reichenbachii var. perbellus (Britton & Rose) Benson 1969d:127.
Echinocereus caespitosus var. perbellus* Weniger 1970:23 nom. inval.
Echinocereus reichenbachii ssp. perbellus* Blum 1997 Seed List nom. nud.
*Echinocereus persolutus  [from Peru and therefor not an Echinocereus]
Echinocereus phaeacanthus* Cactus & Succulent Journal (U.S.)1951:104 nom. nud.
Echinocereus phoeniceus* Lemaire 1868:57 nom. nud. [Probably = Echinocereus coccineus].
Echinocereus phoeniceus* Engelmann ex Rümpler 1885:788 nom. illegit.
  = Cereus phoeniceus Engelmann 1856:284 which was a new name for  Echinocereus
coccineus
Engelman 1848:94.
Echinocereus phoeniceus* var. albispinus Hort. ex Rümpler 1885:789
Echinocereus phoeniceus* var. brevispinus Engelmann ex Schelle 1907:138 nom. nud.
Cereus phoeniceus ssp. conoideus Engelmann 1857:37
  = Echinocereus conoideus.
Echinocereus phoeniceus* var. conoideus Schumann 1896:150
= Echinocereus conoideus.
Echinocereus phoeniceus* var. densus Schelle 1907:138 nom. nud.
Echinocereus phoeniceus* var. inermis Schumann 1896:150
  (see Echinocereus coccineus var. inermis).
Echinocereus phoeniceus* var. longispinus Hort. ex Rümpler 1885:789
Cereus phoeniceus var. pacificus Engelmann ex Orcutt 1886:46
(see Echinocereus pacificus).
Echinocereus phoeniceus* var. rufispinus Hort. ex Rümpler 1885:789
Echinocereus phoeniceus* var. utahensis* Schelle 1907:138 nom. nud.
Echinocereus pleiogonus Croucher 1878:290.
TL: unknown [perhaps an Echinopsis]
Echinocereus pleiogonus* Labouret ex Rümpler 1885:828-829  nom. illegit. (later homonym).
= Cereus pleiogonus Labouret 1853:317.
TL: unknown
Echinocereus plomosus* Köhres Catalog 1984. nom. nud. 
[= Lau 768 from Plomosos, Coahuila ]
Echinocereus polichephalus* Daul 1890:78 nom. nud.
Echinocereus polycephalus
* Hort. Berol. ex Haage 1897 nom. nud.
Echinocereus polyacanthus Engelmann 1848a:104.
TL: Mexico, Chihuahua, Cosihuiriachi, Wislizenus, 1846 (MO).
Synonyms:
Cereus polyacanthus (Engelmann)Engelmann 1849:50.
Echinocereus triglochidiatus var. polyacanthus (Engelmann) Benson 1944:253.
Echinocereus polyacanthus var. acifer* Parker 1996:4 nom. nud. 
  = Echinocereus acifer.
Echinocereus polyacanthus ssp. acifer (Otto ex Salm-Dyck) Taylor 1997:9.
  = Echinocereus acifer.
Echinocereus polyacanthus var. acifer fa. huitcholensis* Parker 1996:4. nom. nud.
= Echinocereus huitcholensis.

Echinocereus polyacanthus var. acifer fa. matthesianus* Parker 1996:4 nom. nud.
= Echinocereus matthesianus*

Echinocereus polyacanthus var. albispinus Hort. ex Schelle 1926:179.
TL: unknown
Echinocereus polyacanthus var. bergeanus Fobe 1911:56.
TL: unknown
Echinocereus polyacanthus var. bergeanus* Hort ex Schelle  1926:179 nom. illegit.
(later homonym).
Echinocereus polyacanthus var. densus (Regel) Taylor 1984:159
= Echinopsis valida var. densa Regel 1852:295.
Note: This name is used by Taylor (1984) as the name of Echinocereus  acifer.
Echinocereus polyacanthus var. durangensis* Parker 1996:4 nom. nud.
= Echinocereus durangensis.
Echinocereus polyacanthus var. durangensis fa. marksianus* Parker 1996:4 nom. nud.
=Echinocereus marksianus.
Echinocereus polyacanthus var. galtieri (Rebut ex Fobe) Schelle 1926:179
= Echinocereus galtieri.
Echinocereus polyacanthus var. huitcholensis (Weber) Taylor 1988:82
= Echinocereus huitcholensis.
Echinocereus polyacanthus ssp. huitcholensis (Weber) Taylor 1997:9
=  Echinocereus huitcholensis.
Echinocereus polyacanthus var. leeanus* Parker 1996:4 nom. nud. 
  = Echinocereus leanus.
Echinocereus polyacanthus var. longispinusHort. ex Schelle 1926:179.
TL: unknown.
Echinocereus polyacanthus var. mombergerianus* Parker 1996:4 nom. nud. 
  = Echinocereus mombergerianus.
Echinocereus polyacanthus ssp. mombergerianus (Frank)Breckwoldt & Matylewicz
1996:15      = Echinocereus mombergerianus.
Echinocereus polyacanthus var. neomexicanus* Weniger 1970:44 nom. inval.
    (no basionym given)     = Echinocereus neomexicanus.
Echinocereus polyacanthus var. nigrispinus Fobe 1911:55.
TL: unknown
Echinocereus polyacanthus var. nigrispinus* Hort. ex Schelle
1926:179 nom. illegit. (later homonym).
Echinocereus polyacanthus var. pacificus (Engelmann ex Orcutt)
Taylor 1984:160     = Echinocereus pacificus.
Echinocereus polyacanthus ssp. pacificus (Engelmann ex Orcutt) Breckwoldt
in Breckwoldt & Matylewicz 1996:15
    = Echinocereus pacificus.
Echinocereus polyacanthus var. rosei* Weniger 1970:43 nom. inval. 
  (no basionym given)     = Echinocereus rosei.
Echinocereus polyacanthus var. rufispinus Fobe 1911:55-56.
TL: unknown
Echinocereus polyacanthus var. rufispinus* Hort. ex Schelle 1926:179 nom. illegit.
   (later homonym).
Echinocereus polycephalus* Blanc 1888:39 nom. nud.
Echinocereus polycephalus* Hort. Berol. ex Haage 1900:160 nom. nud.
Echinocereus polycephalus* var. albispinus* Fric 1924:121 nom. nud.
Echinocereus polycephalus* var. brevinigrispinus* Fric 1924:121
nom. nud.
Echinocereus polycephalus* var. nigrispinus* Fric 1924:121 nom. nud.
Echinocereus polycephalus* var. spinosissimus* Fric 1924:121 nom. nud.
Echinocereus polycephalus var. xphoeniceus* Fric 1924:121 nom. nud.
Echinocereus polypus* Fric 1924:121 nom. nud.
Echinocereus poselgeri Lemaire 1868:57
[A new name needed because of earlier homonym].
= Cereus tuberosus Poselger 1853:131 nom. illegit. 
(later homonym) [not Cereus tuberosus Pfeiffer 1837.]
TL: Texas, near Rio Grande River, above Belleville, Poselger, 
1850 (MO).
Synonyms:
Echinocereus tuberosus* Rümpler 1885: 131 nom. illegit.
          Cereus poselgeri Coulter 1896:398.
Wilcoxia poselgeri (Lemaire) Britton & Rose 1909:434.
Wilcoxia tuberosa*  Berger 1929:134 nom. illegit.
Echinocereus poselgeri fa. australis* Parker 1996:6 nom. nud.
Echinocereus poselgeri ssp. kroenleinii (Cartier) Lange 1995:138.
= Wilcoxia kroenleinii Cartier 1980 (Succulentes 2(2):2-3).
[see Echinocereus kroenleinii].
Echinocereus poselgeri fa. kroenleinii* Parker 1996:6 nom. nud. 
  = Echinocereus kroenleinii.
Echinocereus poselgeri fa. tamaulipensis* Parker 1996:6 nom. nud.
= Echinocereus tamaulipensis.
Echinocereus poselgeri fa. tuberosa* Parker 1996:6 nom. nud.
Echinocereus poselgeri var. waldeisii* Parker 1996:6 nom. nud.
= Echinocereus waldeisii. 
Echinocereus poselgeri var. waldeisii fa. deherdtii* Parker 1996:6 nom. nud. 
   = Echinocereus tamaulipesis var. deherdtii.
Echinocereus poselgerianus Linke 1857:239.
TL: Mexico
Synonyms:
Cereus poselgerianus (Linke) Berger 1905:80.
Echinocereus blanckii var. poselgerianus (Linke) Backeberg 1960:1999.
Note: usually considered a synonym of Echinocereus berlandieri.
Echinocereus primolanatus* Schwarz ex Backeberg 1960:2043 nom.
   inval.  (no type designated).
Echinocereus primolanatus Taylor 1985b:130.
TL: Mexico, Coahuila, Sierra Paila, north of Estacion Marte,
cultivated by D. Parker, 1985(K).
Synonyms:
Echinocereus primolanatus* Schwarz ex Backeberg 1960:2043
        nom. inval. (no type designated).
Echinocereus princeps Föster 1861:163.
TL: unknown [Of unknown origin and perhaps from South America in
which case it would not be an Echinocereus].
Echinocereus procumbens* Lemaire 1868:56 nom. nud.
[Probably = Echinocereus procumbensEngelmann ex Rümpler].
Echinocereus procumbens (Engelmann) Engelmann ex Rümpler 1885:781.
= Cereus procumbens Engelmann 1849:50.
TL: Mexico, Tamaulipas, Burita at mouth of Rio Grande, St. Louis
Volunteers 1846 (MO).
Synonyms:
Echinocereus pentalophus var. procumbens (Engelmann) Fournier 1935:25.
Echinocereus pentalophus fa. procumbens* Parker 1996:4 nom. nud.
        Echinocereus pentalophus ssp. procumbens (Engelmann) Blum & Langein Blum et al.
1998:129. 

Echinocereus procumbens var. gracilior Dautw. ex Schelle 1926:165
TL: unknown
Echinocereus procumbens var. longispinus Hort. ex Hirscht 1902:135.
TL: unknown
Cereus propinquus Salm Dyck ex Otto 1833:366.
  = Echinocereus pentalophus var. simplex.
Echinocereus propinquus De Candolle ex Haage 1892:118
  = Echinocereus pentalophus var. simplex.
Cereus propinquus var. subarticulatus* Pfeiffer ex Förster 1846:373 pro syn.
    = Echinocereus pentalophus var. subarticulatus.
Echinocereus pseudohempelii* Künzler Cat. nom. nud. 
  = Echinocereus kuenzleri.
Echinocereus pseudopectinatus (Taylor) Taylor 1989:74.
= Echinocereus bristolii var. pseudopectinatus Taylor 1985b:120.
TL: Mexico, Sonora, Moctezuma, Lau 607, 1973, cultivated by Taylor, 21 June 1984 (K).
Synonyms:
Echinocereus scopulorum ssp. pseudopectinatus* Blum & Langein Blum et al. 1998:342.
Echinocereus pubellus* Cactus and Succulents Journal (US) 7:5 1936 nom. nud
Echinopsis pulchella var.flore kermesina* Haage ex Förster1846:364. pro syn. 
    = Echinocereus amoenus.
Echinopsis pulchella var. rosea* Labouret 1853:292. pro syn. 
  = Echinocereus amoenus.
Echinocereus pulchellus (Martius) F. Seitz 1870:11.
[see Taylor 1989:75 ].
= Echinocactus pulchellus Martius 1832:342.
TL: Mexico, Hidalgo, Pachuca, Karwinsky (?).
Synonyms:
Cereus pulchellus Pfeiffer 1837:74.
Echinonyctanthus pulchellus (Martius) Lemaire 1839:85.
Echinopsis pulchella (Martius) Zuccarini ex Förster 1846:363.
Note: Usually listed as Echinocereus pulchellus Schumann 1894:185.
Echinocereus pulchellus var. acanthosetus Arias & Guzman in Arias Montes
et al. 1997:63.
TL: Mexico, Oaxaca, between Magdalena Jicotlan & Tepelmeme de
Morelos, Arias, Guzman-Cruz & Gama 958, 29 Oct. 1991 (MEXU).
Synonyms:
Echinocereus pulchellus ssp. acanthosetus (Arias & Guzman)Blum in Blum et al.
         1998:261.
Echinocereus pulchellus ssp. acanthosetus (Arias & Guzman) Blum in
Blum et al. 1998:261. = Echinocereus pulchellus var. acanthosetus
Echinocereus pulchellus var. amoenus (Dietrich) Förster ex Schumann 1897:253
= Echinocereus amoena.
Echinocereus pulchellus ssp. amoenus* Blum 1997 Seed List nom. nud. 
  = Echinocereus. amoenus.
Echinocereus pulchellus var. costatus Halda , Kupčák & Sladk.2000 in Acta Mus. Richnov.
Sect. Nat., 7(1): 34 (2000)
TL: San Luis Potosí/Zacatecas border;  J.J. Halda; J. Sladkovský 8504131 (PR)
Echinocereus pulchellus var. sharpii Taylor 1989:75-77.
TL: Mexico, Nuevo Leon, near San Roberto, 1900m, 24º50´N 100 25´W,
Gonzalez G. 1985 (seed), cult. N. P. Taylor, 4 May 1988 (K).
synonyms:
Echinocereus pulchellus ssp. sharpii (Taylor) Taylor 1997:9
Echinocereus sharpii* S & K Breckwoldt 1998:1. nom. nud.
Echinocereus pulchellus ssp. sharpii (Taylor) Taylor 1997:9
 = Echinocereus pulchellus var. sharpii.
Echinocereus pulchellus subsp. sladkovskyi Halda & Kupčák 2000.
Acta Mus. Richnov. Sect. Nat., 7(1): 34 (2000)
    TL: Hidalgo, MX ; J.J. Halda; J. Sladkovský 850418033 (PR)
Echinocereus pulchellus ssp. venustus Blum & Rischer in Blum et al. 1998:255.
    TL: Mexico San Luis Potosi [Lau 1036] W. Rischer, 5 Dec. 1974 (ZSS A 17984).
Echinocereus pulchellus var. weinbergii (Weingart) Taylor 1985a:272
= Echinocereus weinbergii.
Echinocereus pulchellus ssp. weinbergii (Weingart) Taylor 1997:9
  = Echinocereus weinbergii.
Echinocereus purpureus Lahman 1935:141-143.
TL: Oklahoma, near Medicine Park, Lahman (MO).
Synonyms:
Echinocereus caespitosus var. purpureus* Weniger 1970:23  nom. inval.
Echinocereus reichenbachii var. purpureus* Parker 1996:5 nom. nud.
Echinocereus radians Engelmann 1848a:104.
TL: Mexico, Chihuahua, Cosihuiriachi, Wislizenus, 1846 (MO).
Synonyms:
Cereus adustus var. radians (Engelmann) Engelmann 1849:50.
Echinocereus adustus var. schwarzii fa. radians* Parker 1996:6 nom. nud.
Note: Usually considered a synonym of Echinocereus adustus.
Echinocereus raphicephalus Förster 1861:164.
TL: unknown. [May be of South American origin and not an Echinocereus]
Echinocereus rayonesensis Taylor 1988:75-76.
TL: Mexico, Nuevo Leon, Rayones, Lau 1101 (K).
Echinocereus rayonesensis fa. barkeanus* Parker 1996:3 nom. nud.
[Probably = Echinocereus barkeanus*]
Echinocereus rectispinus Peebles 1938:675.
TL: Arizona, hills near Nogales, 3900 ft., Peebles SF 905
(US 1729266).
Synonyms:
Echinocereus fendleri var. rectispinus (Peebles) Benson 1944:259.
Echinocereus fasciculatus var. rectispinus* Benson 1969d:131 pro syn.
Echinocereus fendleri ssp. rectispinus (Peebles) Taylor 1997:9
Echinocereus rectispinus var. robustus Peebles 1938:675
(see Echinocereus robustus Peebles).
Echinocereus reichenbachianus* Linke 1857:83 nom. nud.
Echinocereus reichenbachianus Hort. ex Haage 1859:20.
[See Echinocereus reichenbachii].
= Echinocactus reichenbachianus (Terscheck) Terscheck ex Fennel
1843:282 (Terscheck, Verzeichnis 3. According to Schumann
1897:271).
Synonyms:
Echinopsis pectinata var. reichenbachiana (Terscheck) Salm-Dyck 1845:26.
Cereus reichenbachianus (Terscheck) Labouret 1853:318.
Echinocereus reichenbachianus* Engelmann ex Haage 1893:23
Note: See note under Echinocereus reichenbachii.
Echinocereus reichenbachianus* Engelmann ex Seitz 1878:11 nom. nud. 
Echinocereus reichenbachianus* Engelmann ex Haage 1893:23
(according to Index Kewensis 2:813) nom. illegit. (later homonym).
Cereus reichenbachianus var. castaneus (Engelmann) Labouret
1853:319 = Echinocereus caespitosus var. castaneus.

Las Cruces, New Mexico 1 May 2006. Permission to copy or reprint
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