Alle Artikel mit dem Schlagwort “Echinocereus Index

Introduction by Terry L. Corbett

Echinocereus is a genus of cactus native to North America.

The index is arranged alphabetically by species name. All names, including names that have never been published under the name Echinocereus are listed alphabetically by species name and then by subspecific name. Autonyms (such as Echinocereus acifer var. acifer) are not listed. For example Cereus dasyacanthus var. spurius will  be found alphabetically under „d“, not „c“, and then by „s“.  To find Echinocereus xxx var. xxx fa. yyy look under Echinocereus xxx fa. yyy. Names in bold are index base names and are followed by information about the basionym and other publication data. For full information of names that are not index base names please check under the name which follows the = sign. Please note that in those cases where the base name is not the basionym, the basionym is listed under the index base name. For the purpose of this index the rank used by the original author has been followed in determining the index base name except in those cases where the original author raised the rank of the plant, the higher rank is followed in determining the index base name. This is for reference purposes only and does not indicate any preference in regard to the proper taxonomic placement of any plant. Synonyms are listed chronologically. Information about the type specimen if available is listed under „Type – locality:“ in the order – location; collector, date (Herbarium).

Bold names are index base names.
*names preceded by an asterisk are excluded from the genus Echinocereus.
names* followed by an asterisk are illegitimate names according to the code of nomenclature.

Echinocereus abbae – Echinocereus crispatus

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Echinocereus Engelmann 1848a:91.
Type: Echinocereus viridiflorus Engelmann

Echinocereus abbeae Parsons 1937:6.
  Type - locality: Mexico, Sonora, Delta Bacuachic River, 30m (29o49'N, 111o51'W); S. H. Parsons, 2 March 1932 (?).
    Nomenclatural synonyms:
      Echinocereus fasciculatus subsp. abbeae* Blum 1997 Seed List. nom. nud.
        Note: Listed erroneously as a synonym of Echinocereus fenderi var. albiflorus (Backeberg 1960:2047, Bravo 1991:70).
          Taylor (1985b:50) treats this species as a taxonomic synonym of Echinocereus fendleri var. fasciculatus.
Echinocereus acifer* Engelmann ex Haage 1859:19 nom. nud. [Probably = Echinocereus acifer Jacobi].
Echinocereus acifer (Otto ex Salm-Dyck) Jacobi 1856:109.
  basionym: Cereus acifer Otto ex Salm-Dyck 180:189.
    Type - locality: unknown; ? (?).
      Nomenclatural synonyms:
        Echinocereus longispinus* Hort. ex Schelle 1926:180 pro syn.
        Echinocereus triglochidiatus var. acifer (Otto ex Salm-Dyck) Bravo 1978:66.
        Echinocereus polyacanthus var. acifer* Parker 1996:4 nom. nud.
        Echinocereus polyacanthus subsp. acifer (Otto ex Salm-Dyck) Taylor 1997:9.
          Note: Taylor (1984:159) uses the name Echinocereus polyacanthus var. densus for this species.
            Cereus acifer* Otto ex Forster 1846:433 nom. nud. is usually listed as a synonym of this species.
Echinocereus acifer* Lemaire 1868:57 nom. nud.
  [Probably = Echinocereus acifer Jacobi].
Echinocereus acifer var. brevispinulus Jacobi 1856:109-110.
  Type - locality: Mexico.
Echinocereus acifer var. brevispinus Haage 1859:19 nom. nud.
  [Probably =  Echinocereus acifer var. brevispinulus Jacobi 1856].
Echinocereus acifer var. diversispinus Schumann 1897:287.
  Type - locality: Mexico, Coahuila & Durango; ? (?).
Echinocereus acifer var. durangensis (Poselger ex Rumpler) Schumann 1897:287.
  = Echinocereus durangensis.
Echinocereus acifer subsp. huitcholensis (Weber) Lange 1994:109.
  = Echinocereus huitcholensis.
Echinocereus acifer var. longispinus* Haage 1892:116 nom. nud.
Echinocereus acifer var. tenuispinus Jacobi 1856:109.
  Type - locality: North America.
Echinocereus acifer subsp. topiensis (Rischer & Trocha) Rischer 2004:88.
  = Echinocereus topiensis W. Rischer & Trocha 1999.
Echinocereus acifer var. trichacanthus Hort. ex Hildmann 1891:44.
  Type - locality: unknown; ? (?).
Echinocereus acifer subsp. tubiflorus Rischer in Blum et al. 1998a:1. [see Blum et al. 1998b:409-410].
  Type - locality: Mexico, Zacactecas - Durango border, San Juan Capistran; Rischer, 17 April 1990 (ZSS AX 17983).
Echinocereus acifer subsp. ventanensis Rischer 2009: 63.
  Type - locality: Mexiko: Sinaloa, 63.4 miles west of El Salto (Durango), 6300 alt.; Kimnach 563, 25. Nov. 1964 (HNT 8399).
Cereus acinaciformis* Steudel 1841:333 pro syn.
  = Echinocereus cinerascens.
Cereus acinaciformis* Hort. Berol. ex Salm-Dyck 1845:27 pro syn.
  = Echinocereus cinerascens.
Cereus aciniformis* Otto 1833:336 nom. nud.
  = Echinocereus cinerascens var. crassior.
Cereus aciniformis* Hort. Berol. ex Pfeiffer 1837:101 pro syn.
  = Echinocereus cinerascens var. crassior.
Echinocereus adustus Engelmann 1848a:104.
  Type - locality: Mexico, Chihuahua, Cosihuiriachi, 1900 m.; Wislizenus, 1846. (MO 2371547).
    Nomenclatural synonyms:
      Cereus adustus (Engelmann) Engelmann 1849:50.
      Echinocereus pectinatus var. adustus (Engelmann) Daul 1890:78.
      Echinocereus pectinatus fa. adustus (Engelmann) Voss 1894:378.
      Echinocereus pectinatus var. adustus* Schumann 1897:271.
      Echinocereus caespitosus var. adustus (Engelmann) Engelmann ex Borg 1937:174.
        Note: Echinocereus radians and Echinocereus rufispinus are usually considered taxonomic synonyms of this species.
Echinocereus adustus var. bonatzii (Roemer) Taylor 1998:15.
  = Echinocereus bonatzii Roemer 1995:10.
Cereus adustus var. radians (Engelmann) Engelmann 1849:50
  = Echinocereus radians.
Echinocereus adustus subsp. roemerianus W. Rischer 2003a:44.
  Type - locality: Mexico, Durango, Canatlan; R. C. Roemer 6066714 (UNAM).
Echinocereus adustus var. schwarzii (Lau) Taylor 1985a:268.
  = Echinocereus schwarzii.
Echinocereus adustus subsp. schwarzii (Lau) Taylor 1997:8.
  = Echinocereus schwarzii.
Echinocereus adustus var. schwarzii fa. madrensis* Parker 1996:6. nom. nud.
  = Echinocereus madrensis.
Echinocereus adustus var. schwarzii fa. radians* Parker 1996:6. nom. nud.
  = Echinocereus radians.
Echinocereus adustus var. schwarzii fa. rufispinus* Parker 1996:6. nom. nud.
  = Echinocereus rufispinus.
Echinocereus aggregatus Rydberg 1906:146.
  = Mammillaria aggregata* Engelmann 1848b:157. nom. prov.
  = Mammillaria aggregata Engelmann ex Watson 1878:398 pro syn.
  [Rydberg used this name instead of Echinocereus coccineus].  see Echinocereus coccineus subsp. aggregatus.
Echinocereus aguirrei* Glass 1965:160 nom. nud. [a form of Echinocereus pulchellus].
Echinocereus aguirreanus* Hort. ex Blum et al. 1998b:253 pro syn.
  = Echinocereus pulchellus.
Echinocereus albatus* Backeberg 1960:2007-2008 nom. inval. (no type designated).
  Nomenclatural synonyms:
    Echinocereus albispinus* Hort. ex Backeberg 1960:2007 pro syn.
    Echinocereus longisetus var. albatus* Sterk 1980:12-13 nom. inval.
      Note: Usually considered a taxonomic synonym of Echinocereus nivosus.
Echinocereus albiflorus Weingart 1933:156.
  Type - locality: Mexico, Chihuahua; Halbinger (?).
    Nomenclatural synonyms:
      Echinocereus fendleri var. albiflorus (Weingart)  Backeberg 1960:2047.
      Echinocereus fendleri fa. albiflorus* Parker 1996: 2 nom. nud.
        Note: Backeberg 1960:2047 and Bravo 1991:70 erroneously list Echinocereus abbeae as a synonym of this species.
          Taylor (1985b) lists this species as a taxonomic synonym of Echinocereus fendleri. It is probably a form of E. hempelii.
Echinocereus albiflorus* Lowry 1937:20 nom. nud. [Probably a white flowering form of Echinocereus enneacanthus].
Echinocereus albispinus Lahman 1935:143.
  Type - locality: Oklahoma, Medicine Park; Lahman (MO).
    Nomenclatural synonyms:
      Echinocereus baileyi var. albispinus (Lahman) Backeberg 1941:4.
      Echinocereus longispinus var. albispinus* Backeberg 1960:2010.
      Echinocereus reichenbachii var. albispinus (Lahman) Benson 1969b:127.
      Echinocereus reichenbachii var. baileyi fa. albispinus* Mesa Garden Seed Catalog 1996 nom. nud.
        Note: The name Echinocereus reichenbachii var. albispinus is used for  Echinocereus baileyi by Benson (1969b:127).
          Taylor (1985b) lists this species as a taxonomic synonym of Echinocereus reichenbachii var. baileyi.
Echinocereus albispinus* Hort. ex Backeberg 1960:2007 pro syn. = Echinocereus albatus.
Echinocereus amoenus (Dietrich) Weber ex Haage 1892:116.
  basionym: Echinopsis amoena Dietrich 1844:187-188.
    Type - locality: Mexico, Hidalgo, near San Mateo 7500 ft.; Ehrenberg (?).
      Nomenclatural synonyms:
        Echinopsis pulchella var. flore kermesina* Haage ex Forster 1846:364. pro syn.
        Echinopsis pulchella var. amoena (Dietrich) Forster 1846:364.
        Echinopsis pulchella var. rosea* Labouret 1853:292. pro syn.
        Cereus amoenus (Dietrich) Hemsley 1880:540.
        Echinocereus pulchellus var. amoenus (Dietrich) Forster ex  Schumann 1897:253.
        Echinocereus pulchellus subsp. amoenus (Dietrich) Blum 1997 Seed list. nom. nud.
        Note: Echinocereus amoenus is usually attributed to Schumann 1894:185.  Taylor (1989:75) notes that the plant
          usually known in cultivation as "Echinocereus amoenus" is instead a form of Echinocereus weinbergii.
Echinocereus amoenus var. aguirre* Zehnter & Kunzler Catalog nom. nud. = Echinocereus aguirrei*.
Echinocereus amoenus var. albiflorus* Hort. ex Blum et al. 1998b:259 pro syn.
Echinocereus angusticeps Clover 1935:79.
  Type - locality: Texas, Hidalgo Co., Linn; Clover 15261 (MICH).
    Nomenclatural synonyms:
      Echinocereus papillosus var. angusticeps (Clover) Marshall & Bock 1941:119.
      Echinocereus blanckii var. angusticeps (Clover) Benson 1969b:260.
      Echinocereus berlandieri var. angusticeps (Clover) Benson 1976:59.
      Echinocereus papillosus subsp. angusticeps* Blum 1997 Seed List nom. nud.
      Note: Benson uses the names Echinocereus blanckii var. angusticeps and  Echinocereus berlandieri var. angusticeps for
        Echinocereus papillosus subsp. papillosus.
Echinocereus anneacanthus* Daul 1890:73. (misprint for E. enneacanthus).
Echinocereus apachensis Blum & Rutow in Blum et al. 1998a:1. [see Blum et al. 1998b:101.]
  Type - locality: Arizona, Maricopa Co. Fish Creek; Mc Kelvey 735, 18 Feb. 1929  (US 1532949).
    Nomenclatural synonyms:
      Echinocereus bonkerae subsp. apachensis (W.Blum & Rutow) A.D. Zimmerman in Baker et al. 2009:120.
Echinocereus arizonicus Rose ex Orcutt 1926:3.
  Type - locality: Arizona, Boundary Monument between Pinal & Gila Counties, 4700 ft.; C. R. Orcutt, July 1922 (NY).
    Nomenclatural synonyms:
      Echinocereus triglochidiatus var. arizonicus (Rose ex Orcutt) Benson 1969a:21.
      Echinocereus coccineus var. arizonicus (Rose ex Orcutt)  Ferguson 1989:221.
Echinocereus arizonicus subsp. matudae (Bravo) Rutow in Lange & Rutow 1994:51.
  = Echinocereus matudae.
Echinocereus arizonicus subsp. nigrihorridispinus Blum & Rutow in Blum et al. 1998a:3. [see Blum et al. 1998b:370-371].
  Type - locality: New Mexico, Luna Co.; HK 1054 (ZSS AX 17982).
Echinocereus armatus (Poselger) Poselger ex Backeberg & Knuth 1935:318.
  = Echinocereus pectinatus var. armatus.
*Echinocereus arrojadei Lofgren [from Brazil and thus not an Echinocereus].
Echinocereus australis* Backeberg 1937:21 nom. nud.
Echinocereus australis* Hort. ex Backeberg 1960:2045 pro syn.
  = Echinocereus fendleri.
Echinocereus bakeri W.BLUM, OLDACH & J.OLDACH, Int. Cact. Advent. 106-107: 22. 2015.
  Type - locality: Arizona, Yavapai County, 6 km. north of Clarkdale ; Marc A. Baker 13930, 18 March 2001 (ASU 0076433)
Echinocereus baileyi Rose 1909b:403.
  Type - locality: Oklahoma, Wichita Mountains.; V. Bailey, Aug. 1906 (US 53167).
    Nomenclatural synonyms:
      Echinocereus reichenbachii var. baileyi (Rose) Taylor 1985b:133.
      Echinocereus reichenbachii subsp. baileyi (Rose) Taylor 1997:9.
        Note: The name Echinocereus reichenbachii var. albispinus is used for this species by Benson (1969b:127 ).
Echinocereus baileyi var. albispinus (Lahman) Backeberg 1941:4.
  = Echinocereus albispinus Lahman.
Echinocereus baileyi var. brunispinus Backeberg 1941:4.
  Type - locality: Oklahoma, Wichita Mountains; ? (?).
    Note: Backeberg 1960:2010 lists Echinocereus longispinus and Echinocereus oklahomensis as synonyms of this variety.
Echinocereus baileyi var. caespiticus* Backeberg 1960:2011 nom. inval. (no type designated).
Echinocereus baileyi var. flavidispinus Backeberg 1941:4.
  Type - locality: Oklahoma, Wichita Mountains; ? (?).
Echinocereus baileyi var. roseispinus Backeberg 1941:4.
  Type - locality: Oklahoma, Wichita Mountains; ? (?)
Echinocereus barcelona* Orcutt 1926:4 nom. nud.
Echinocereus barcena* Rebut ex Hirscht 1896:126 nom. nud.
Echinocereus barcena Fobe 1911:56.
  Type - locality: unknown; ? (?) [from central Mexico according to Borg 1937:178].
    Nomenclatural synonyms:
      Echinocereus parkeri fa. barcena* Parker 1996:3 nom. nud.
Echinocereus barcena* Rebut ex Berger 1929 nom. illegit. (later homonym) [Perhaps  = Echinocereus barcena Fobe].
Echinocereus barkeanus* Heyer 1989:25 nom. nud. [= Lau 1101].
  = Echinocereus rayonesensis.
Echinocereus barthelowanus Britton & Rose 1922:41.
  Type - locality: Baja California, Isla Magdalena, Santa Maria Bay; J. N. Rose, 18 March 1911 (US 16278).
Echinocereus bauerii Rother 1902:61.
  Type - locality: ? (?).
    Nomenclatural synonym:
      Echinocereus pectinatus var. bauerii Rother 1923:127.
Echinocereus bergeanus* Haage 1900:156 nom. nud.
Echinocereus berlandianus* Orcutt 1926:3 [error in spelling for Echinocereus berlandieri].
Echinocereus berlandieri (Engelmann) Haage 1859:19.
  basionym: Cereus berlandieri Engelmann 1856:286. [see Engelmann 1859:38, t.58].
    Type - locality: Texas, Nueces River; Berlandier 2423, April 1834 (MO).
      Nomenclatural synonyms:
        Echinocereus blankii* Hort. ex Palmer 1865:92 pro syn.
        Echinocereus blanckii var. berlandieri (Engelmann) Backeberg 1960:1999.
          Note: Echinocereus poselgerianus is usually considered a taxonomic synonym of this species. This species is
            often erroneously treated as a synonym of Echinocereus blanckii.
Echinocereus berlandieri var. angusticeps (Clover) Benson 1976:59.
  = Echinocereus angusticeps.
Echinocereus berlandieri var. blanckii* Fournier 1935:21 nom. illegit.
  = Echinocereus blanckii.
Echinocereus berlandieri var. longispinus* Hort. ex Schelle 1926: 163 nom. nud.
Echinocereus berlandieri var. papillosus (Linke ex Rumpler) Benson 1976:59.
  = Echinocereus papillosus.
Echinocereus berlandieri var. poselgerianus (Linke) Lode 1992:17.
  = Echinocereus poselgerianus.
*Echinocereus bertinii = Austrocactus bertinii.
*Echinocereus bicolor Galeotti  [error in spelling for Echinocactus bicolor].
Cereus bigelovii* Engelmann 1857: t. 4 nom. nud.
  = Echinocereus mojavensis.
Cereus bigelovii var. zuniensis* Engelmann 1857: t. 4 nom. nud.
  = Echinocereus mojavensis var. zuniensis.
Echinocereus blanckii (Poselger) Poselger ex Rumpler 1886:779.
  basionym: Cereus blanckii Poselger 1853:134.
    Type - locality: Mexico, Tamaulipas, near Camargo; Poselger (?).
      Nomenclatural synonyms:
        Echinocereus berlandieri var. blanckii* Fournier 1935:21 nom. illegit.
          Note: This species is usually considered a taxonomic synonym of Echinocereus enneacanthus subsp. brevispinus. The
            name Echinocereus blanckii has been errroneously applied to Echinocereus berlandieri.
Echinocereus blanckii var. angusticeps (Clover) Benson 1969b:260.
  = Echinocereus angusticeps.
    Note: This is the name Benson uses for Echinocereus papillosus subsp. papillosus.
Echinocereus blanckii var. berlandieri (Engelmann) Backeberg 1960:1999.
  = Echinocereus berlandieri.
Echinocereus blanckii var. inermis* Heyer 1989:26 pro syn.
  = Echinocereus moricallii.
Echinocereus blanckii var. leonensis (Mathsson) Backeberg 1960:1999.
  = Echinocereus leonensis.
Echinocereus blanckii var. papillosus (Linke ex Rumpler) Benson 1969d:126.
  = Echinocereus papillosus.
Echinocereus blanckii var. poselgerianus (Linke) Backeberg 1960:1999.
  = Echinocereus poselgerianus.
Echinocereus blankii* Hort. ex Palmer 1865:92 pro syn.
  = Echinocereus berlandieri.
Echinocereus blumii D. Felix & H. Bauer in Blum, W., D. Felix & H. Bauer. 2012: 220 - 237.
  Type - locality: Mexiko: Coahuila: Municipio Acuna, El Conejo, 1200 m.; det. W. Blum 2006 (KR 21304).
Echinocereus bolansis* Runge ex Haage 1892:117 pro syn.
  = Echinocereus conglomeratus.
*Echinocereus boliviensis Poselger ex Schumann 1897:290.[from Bolivia which means it is not an Echinocereus].
Echinocereus bonatzii Romer 1995:98-113.
  Type - locality: Durango, Sierra Mezquital; H. J. Bonatz 12388 (ZSS).
    Nomenclatural synonyms:
      Echinocereus pamanesiorum subsp. bonatzii (Romer) Romer 1997:45.
Echinocereus bonkerae Thornber & Bonker 1932:71-73, 85.
  Type - locality: Arizona, Pinal Mountains near Oracle; Thornber & Bonker (ARIZ 156240).
    Nomenclatural synonyms:
      Echinocereus fendleri var. bonkerae (Thornber & Bonker) Benson 1944:260.
      Echinocereus boyce-thompsonii var. bonkerae (Thornber & Bonker) Peebles 1949:191-192 .
      Echinocereus fasciculatus var. bonkerae (Thornber & Bonker) Benson 1969a:21.
      Echinocereus engelmannii var. bonkerae* Mesa Garden Seed Catalog 1996. nom. nud.
      Echinocereus fasciculatus subsp. bonkerae (Thornber & Bonker) Taylor 1997:8.
Echinocereus bonkerae subsp. apachensis (W.Blum & Rutow) A.D. Zimmerman in Baker et al. 2009:120.
  = Echinocereus apachensis.
Echinocereus boyce-thompsonii Orcutt 1926:4.
  Type - locality: Arizona, Boyce-Thompson Arboretum; Orcutt (?).
    Nomenclatural synonyms:
      Echinocereus fendleri var. boyce-thompsonii (Orcutt) Benson 1944:260.
      Echinocereus fasciculatus var. boyce-thompsonii (Orcutt) Benson 1969a:22.
      Echinocereus engelmannii var. boyce-thompsonii* Mesa Garden Seed Catalog 1996 nom. nud.
      Echinocereus fasciculatus subsp. boyce-thompsonii (Orcutt) Taylor 1997:8.
Echinocereus boyce-thompsonii var. bonkerae (Thornber & Bonker) Peebles 1949:191-192.
  = Echinocereus bonkerae.
Echinocereus brandegeei (Coulter) Coulter ex Schumann 1897:290.
  basionym: Cereus brandegeei Coulter 1896:389.
    Type - locality: Baja California, between El Campo Aleman & San Gregorio; T.S. Brandegee, 1889 (UC).
      Note: Echinocereus mamillatus and Echinocereus sanborgianus are usually considered taxonomic synonyms of this species.
Echinocereus brandegeei fa. mamillatus* Parker 1996:1 nom. nud.
  = Echinocereus mamillatus.
Echinocereus brandegeei var. sanborgianus* Lode 1994: no. 1272 nom. inval.
  = Echinocereus sanborgianus.
Echinocereus brandegeei fa. sanborgianus* Parker 1996:1 nom. nud.
  = Echinocereus sanborgianus.
Echinocereus brenbergii* Lowry 1933:410 nom. nud.
Echinocereus bristolii Marshall 1938:160-161.
  Type - locality: Sonora, Soyopa, 5000 ft.; Bristol & Marshall, 1934 (DS 251175).
    Nomenclatural synonyms:
      Echinocereus pectinatus var. bristolii (Marshall) Marshall 1956b:81.
Echinocereus bristolii subsp. floresii (Backeberg) Blum & Lange in Blum et al. 1998a:3. [see Blum et al. 1998b:353].
  = Echinocereus floresii.
Echinocereus bristolii var. pseudopectinatus Taylor 1985b:120.
  see Echinocereus pseudopectinatus.
Echinocereus caespitosus (Engelmann) Engelmann 1848a:110.
  basionym: Cereus caespitosus Engelmann 1845:247.
    Type - locality: Texas, Austin Co., Near Cat Springs; Lindheimer, June 1845 (MO).
      Nomenclatural synonyms:
        Cereus caespitosus var. minor* Engelmann 1856:280 [Based on same type as Echinocereus caespitosus].
        Echinocereus pectinatus var. caespitosus (Engelmann) Schumann 1897:272.
        Echinocereus caespitosus var. minor* Engelmann ex Weniger 1970:22 nom. inval.
        Echinocereus reichenbachii fa. caespitosus* Parker 1996:5 nom. nud.
        Echinocereus reichenbachii var. caespitosus* Mesa Garden Seed Catalog 1996 nom. nud.
        Echinocereus reichenbachii subsp. caespitosus (Engelmann) Blum & Lange in Blum et al.1998a:3  [see Blum et al.
          1998b:324].
          Note: Echinocereus caespitosus is often treated as a taxonomic synonym of Echinocereus reichenbachii.
Echinocereus caespitosus var. adustus (Engelmann) Engelmann ex Borg 1937:174.
  = Echinocereus adustus.
Echinocereus caespitosus var. armatus (Poselger) Poselger ex Borg 1937:174.
  = Echinocereus pectinatus var. armatus.
Echinocereus caespitosus fa. armatus subfa. castaneas (Engelmann) Schelle 1907:131.
  = Echinocereus caespitosus var. castaneus.
Echinocereus caespitosus var. candicans* Hort. ex Borg 1937:174 pro syn.
  = Echinocereus reichenbachii.
Echinocereus caespitosus var. castaneus (Engelmann) Engelmann ex Rumpler 1886:811.
  basionym: Cereus caespitosus var. castaneus Engelmann 1850:203.
    Type - locality: Texas, Granitic region of the Llano; Lindheimer, 1847 (MO).
      Nomenclatural synonyms:
        Cereus reichenbachii var. castaneus (Engelmann) Labouret 1853:319.
        Echinocereus pectinatus var. castaneus (Engelmann) Daul 1890:78.
        Echinocereus castaneus (Engelmann) Orcutt 1926:3.
        Echinocereus pectinatus var. jamauvei *Frič ex Orcutt 1926:4 pro syn.
        Echinocereus pectinatus var. adustus fa. castaneus (Engelmann) Engelmann ex Schelle 1926:174.
        Echinocereus caespitosus fa.. castaneus Engelmann ex Borg 1937:174.
        Echinocereus pectinatus fa. castaneus (Engelmann) Krainz 1967:50.
          Note: This variety is now usually considered a taxonomic synonym of  E. caespitosus.
Echinocereus caespitosus fa. castaneus Engelmann ex Borg 1937:174.
Echinocereus caespitosus var. chrysacanthus (Schumann) Schumann ex Borg 1937:174.
  = Echinocereus pectinatus var. chrysacanthus.
Echinocereus caespitosus var. lanugunosus* Walton 1898:12 nom. nud.
Echinocereus caespitosus var. major (Engelmann) Engelmann ex Rumpler 1886:811.
  basionym: Cereus caespitosus var. major Engelmann 1856:280.
    Type - locality: Texas, New Braunfels; Lindheimer, 1851 (MO).
      Note: This variety is now usually considered a taxonomic synonym of  E. caespitosus.
Echinocereus caespitosus var. minor* Engelmann ex Weniger 1970:22 nom. inval.
  basionym: Cereus caespitosus var. minor* Engelmann 1856:280. new name for Echinocereus caespitosus var. caespitosus.
Echinocereus caespitosus var. perbellus* Weniger 1970:23 nom. inval. (no basionym given).
  = Echinocereus perbellus.
Echinocereus caespitosus var. purpureus* Weniger 1970:23 nom. inval. (no basionym given).
  = Echinocereus purpureus.
Echinocereus caespitosus var. reichenbachii* Ters. ex Borg 1937:174 nom. illegit.
  = Echinocereus reichenbachii.
Echinocereus caespitosus var. rubescens* Hort. ex Borg 1951:221 nom. inval. (no Latin diagnosis).
Echinocereus caespitosus var. rufispinus (Schumann) Schumann ex Borg 1951:221.
  = Echinocereus pectinatus var. rufispinus.
Echinocereus caespitosus var. tamaulipensis (Frič) Frič ex Borg 1937:174.
  = Echinocereus pectinatus var. tamaulipensis.
*Echinocereus candicans Rumpler 1886 = Echinopsis candicans.
*Echinocereus candicans var. tenuispinus = Echinopsis candicans
Echinocereus candicans* Runge ex Haage 1859:19 pro syn.
  = Echinocereus pectinatus var. rigidissimus.
Echinocereus candicans var. couranti* Walton 1898:12. nom. nud.
Echinocereus candicans var. dumisnilianus* Walton 1898:12. nom. nud.
Echinocereus candicans var. gladiatus* Walton 1898:12. nom. nud.
Echinocereus candicans var. robustus* Walton 1898:12. nom. nud.
Echinocereus canyonensis Clover & Jotter 1941:417-418.
  Type - locality: Arizona, Coconino Co., Bass Cable, Grande Canyon; Clover & Jotter 2317 (MICH).
    Nomenclatural synonyms:
      Echinocereus coccineus subsp. canyonensis* Blum 1997 Seed List nom. nud.
        Note: Taylor (1985b) lists this species as a synonym of Echinocereus triglochidiatus var. melanacanthus, which currently
          has the proper name Echinocereus coccineus.
Echinocereus canus (Powell & Weedin) Felix & Bauer in Blum et al. 2012:258 - 276.
  = Echinocereus viridiflorus var. canus Powell & Weedin 2004:273.
Echinocereus carmenensis Blum, Lange & Scherer in Blum et al. 1998a:3. [see Blum et al. 1998b:221].
  Type - locality: Mexico, Coahuila, Sierra del Carmen; SB 1476 (ZSS A 17986).
Echinocereus carnosus Rumpler 1886:796-797.
  Type - locality: Texas; ? (?).
    Nomenclatural synonyms:
      Echinocereus enneacanthus fa. carnosus (Rumpler) Schelle 1907:127.
      Echinocereus enneacanthus var. carnosus (Rumpler) Quehl 1908:114.
      Echinocereus enneacanthus var. brevispinus fa. carnosus* Parker 1996:2 nom. nud.
        Note: This species is usually considered a taxonomic synonym of Echinocereus enneacanthus subsp. brevispinus.
Echinocereus castaneus* Engelmann ex Haage 1859:19 nom. nud.
Echinocereus castaneus (Engelmann) Orcutt 1926:3.
  = Echinocereus caespitosus var. castaneus.
*Echinocereus centralis Rose = Echinomastus centralis.
Echinocereus cereiformis* Auct. ? (Kakteenkunde 1935:1) nom. inval. (no Latin diagnosis).
  Note: Taylor (1985b) lists this name as a synonym of Echinocereus berlandieri.
Echinocereus cernosus* Frič 1929:24 nom. nud. [Probably = Echinocereus pectinatus].
Echinocereus chaletii Rischer 2009a:39-57
  Type - locality: Mexico, Chihuahua, Municipio Guachochic, Cumbres de Guarachi; M. Chalet 4335 (IBUG 179029).
Echinocereus chihuahuensis* Shields 1961:4. nom. inval. (no Latin diagnosis).
*Echinocereus chiloensis = Echinopsis chiloensis.
Echinocereus chisoensis Marshall 1940:15.
  Type - locality: Texas, Chisos Mtns.; F. Radley, 10 Apr. 1939 (DS 263216).
    Nomenclatural synonyms:
      Echinocereus reichenbachii var. chisoensis (Marshall) Benson 1969d:127.
Echinocereus chisoensis var. fobeanus (Oehme) Taylor 1985a:261
  = Echinocereus fobeanus.
Echinocereus chisosensis subsp. fobeanus (Oehme) Taylor 2005:10.
  = Echinocereus fobeanus.
Echinocereus chisoensis var. metornii* Parker 1996:5 nom. nud.
  = Echinocereus metornii.
Echinocereus chloranthus (Engelmann) Engelmann ex Haage 1859:19.
  basionym: Cereus chloranthus Engelmann 1856:278-279. [see Engelmann 1859:29].
    Type - locality: Texas, stony hills near Frontera [El Paso]; Wright 95, 2 Apr. 1852 (MO 2016809).
      Nomenclatural synonyms:
        Echinocereus viridiflorus var. chloranthus (Engelmann) Backeberg 1960:2015.
        Echinocereus viridiflorus fa. chloranthus (Engelmann) Krainz 1967:51.
        Echinocereus viridiflorus subsp. chloranthus (Engelmann) Taylor 1997:10.
          Note: Usually listed as Echinocereus chloranthus Engelmann ex Rumpler 1886:814.
Echinocereus chloranthus var. albispinus* Frič 1924:121 nom. nud.
Echinocereus chloranthus var. cylindricus (Engelmann) Taylor 1984:169.
  = Echinocereus viridiflorus var. cylindricus.
Echinocereus chloranthus subsp. cylindricus (Engelmann) Blum & Lange in Blum et al. 1998a:3. [see Blum et al. 1998b: 205].
  = Echinocereus viridiflorus var. cylindricus.
Echinocereus chloranthus var. davisii* Cullmann 1984:pl. 154.  nom. inval.
  = Echinocereus davisii.
Echinocereus chloranthus var. flaviflorus* Hort. nom. nud.
Echinocereus chloranthus var. flavispininus* Ito 1952:133 nom. inval. (no Latin diagnosis).
Echinocereus chloranthus var. livermorensis* Kolle 1978:34 nom. inval.
  = Echinocereus russanthus subsp.. weedinii.
Echinocereus chloranthus var. longisetus* Hort. nom. nud.
  = Echinocereus russanthus subsp.. weedinii.
Echinocereus chloranthus var. milleri* Hochstatter Seed List 1997 nom. nud.
  [See Echinocereus milleri*].
Echinocereus chloranthus var. neocapillus Weniger 1969:39-41.
  Type - locality: Texas, 5 to 10 miles south of Marathon; Weniger 711 (UNM).
    Nomenclatural synonyms:
      Echinocereus viridiflorus var. neocapillus (Weniger) Leuck ex Zimmerman in Powell & Weedin 2004:267.
      Echinocereus russanthus var. neocapillus* Mesa Garden Seed Catalog 1996 nom. nud.
      Echinocereus neocapillus Blum & Lange in Blum et al. 1998b:219.
Echinocereus chloranthus subsp. rhyolithensis Blum & Lange in Blum et al. 1998a:4. [see Blum et al. 1998b:208.]
  Type - locality: New Mexico, Sierra County, Lake Valley; Pierce 110, 1961 (UNM).
Echinocereus chloranthus var. roseiflorus* Ito 1981.(according to Evanich 1989:103) nom. nud.
Echinocereus chloranthus var. russanthus (Weniger) Lamb ex Rowley 1974a:7
  = Echinocereus russanthus.
Echinocereus chloranthus var. senilis* Frič 1924:121 nom. nud.
Echinocereus chloranthus var. vulpis-cauda* Chudleigh 1967:43 nom. nud.
  [Probably = Echinocereus russanthus].
Echinocereus chloranthus subsp. weedinii* Hochstatter Seed List 1997 nom. nud.
  = Echinocereus russanthus subsp. weedinii.
Echinocereus chlorophthalmus (Hooker) Britton & Rose 1913:242.
  basionym: Echinocactus chlorophthalmus Hooker 1848:t. 4373.
    Type - locality: Mexico, Hidalgo, Real del Monte; ? (?).
      Nomenclatural synonyms:
        Echinocereus cinerascens fa. chlorophthalmus* Parker 1996:2 nom. nud.
        Note: This species is often considered a taxonomic synonym of Echinocereus cinerascens.
Echinocereus chrysocentrus (Engelmann & Bigelow) Orcutt 1926:4.
  = Echinocereus engelmannii var. chrysocentrus.
Echinocereus cinerascens* Engelmann ex Haage 1859:19 nom. nud.
  [Probably = Echinocereus cinerascens].
Echinocereus cinerascens* Lemaire 1868:56 nom. nud.
  [Probably = Echinocereus cinerascens].
Echinocereus cinerascens (De Candolle) De Candolle ex Rumpler 1886:786.
  basionym: Cereus cinerascens De Candolle 1828:116.
    Type - locality: Mexico, Zimapan; Coulter 23 (?).
      Nomenclatural synonyms:
        Cereus acinaciformis* Steudel 1841:333 pro syn.
        Cereus acinaciformis* Hort. Berol. ex Salm-Dyck 1845:27 pro syn.
        Echinocereus deppei*(Salm-Dyck) Schumann 1895:123 pro syn.
        Echinocereus undulatus* Hildmann ex Schumann 1895:124 pro syn.
          Note: Cereus deppei, Echinocereus cirrhiferus and Echinocereus glycimorphus are usually considered taxonomic synonyms
            of  Echinocereus cinerascens.
Echinocereus cinerascens var. caesius* Hort. ex Schelle 1926:169 nom. nud.
Echinocereus cinerascens fa. chlorophthalmus* Parker 1996:2 nom. nud.
  = Echinocereus chlorophtahlmus.
Echinocereus cinerascens var. crassior (De Candolle) De Candolle ex Rumpler 1886:786-787.
  basionym: Cereus cinerascens var. crassior De Candolle 1828:116.
    Type - locality: Mexico, Zimapan; Coulter (?).
      Nomenclatural synonyms:
        Cereus aciniformis* Otto 1833:336 nom. nud.
        Cereus aciniformis* Hort. Berol. ex Pfeiffer 1837:101 pro syn.
Echinocereus cinerascens var. ehrenbergii (Pfeiffer) Bravo 1974:47.
  = Echinocereus ehrenbergii.
Echinocereus cinerascens subsp. ehrenbergii (Pfeiffer) Blum & Rutow in Blum et al. 1998a:4. [see Blum et al. 1998b:140-141].
  = Echinocereus ehrenbergii.
Cereus cinerascens var. fendleri (Engelmann) Bois 1928:371.
  = Echinocereus fendleri.
Echinocereus cinerascens fa. glycimorphus* Parker 1996:2 nom. nud.
  = Echinocereus glycimorphus.
Echinocereus cinerascens var. schmidtii* Haage 1859:19 nom. nud.
Echinocereus cinerascens var. septentrionalis Taylor 1988:68-69.
  Type - locality: Mexico, San Luis Potosi, Entronque Huizache; Taylor 281, 10 July 1986 (MEXU).
    Nomenclatural synonyms:
      Echinocereus cinerascens subsp. septentrionalis.
Echinocereus cinerascens subsp. septentrionalis Taylor 1997:8.
  = Echinocereus cinerascens var. septentrionalis .
Echinocereus cinerascens fa. spinibarbis* Parker 1996:2 nom. nud.
  = Echinocereus spinibarbis* Hildmann [not Salm-Dyck].
Echinocereus cinerascens var. tenuior (De Candolle) De Candolle ex Rumpler 1886:776.
  basionym: Cereus cinerascens var. tenuior De Candolle 1828:116.
  Type - locality: Mexico, Zimapan; Coulter (?).
Echinocereus cinerascens var. tulensis Taylor 1988:69.
  = Echinocereus tulensis.
Echinocereus cinerascens subsp. tulensis (Bravo) Taylor 1997:8.
  = Echinocereus tulensis.
*Echinocereus cinnabarinus = Echinopsis cinnabarina.
Echinocereus cirrhiferus* Engelmann ex Haage 1859:19 nom. nud.
Echinocereus cirrhiferus (Labouret) Labouret ex Rumpler 1886:778.
  Basionym: Cereus cirrhiferus Labouret 1853:311.
    Type - locality: Mexico; ? (?).
      Note: Usually considered a taxonomic synonym of Echinocereus cinerascens.
Echinocereus cirrhiferus var. virescens* Walton 1898:12 nom. nud.
*Echinocereus clavatus = Eridisia spiniflora.
Echinocereus claviformis* Haage 1859:19. nom. nud.
Echinocereus claviformis (Regel & Klein) Haage 1893:22.
  basionym: Cereus claviformis Regel & Klein 1860:46.
    Type - locality: Mexico, Karwinsky ( ? ).
Echinocereus coccineus Engelmann 1848a:93.
  Type - locality: New Mexico, Wolf Creek, Wislizenus, 24 June 1846 (MO 2016699).
    Nomenclatural synonyms:
      Cereus coccineus (Engelmann) Engelmann 1849:51 nom. illegit. [not Cereus coccineus Salm- Dyck 1828].
      Cereus phoeniceus* Engelmann 1856:284. new name for Echinocereus coccineus.
      Echinocereus phoeniceus* Engelmann ex Rumpler 1886:788 nom. illegit.
      Echinocereus melanacanthus* Engelmann ex Earl 1963:61 printing error.
      Echinocereus triglochidiatus var. coccineus (Engelmann) Engelmann ex Marshall & Bock 1941:117.
        Note: Benson uses the name Echinocereus triglochidiatus var. melanacanthus for this species.
Echinocereus coccineus subsp. aggregatus Blum, Lange & Rutow in Blum et al. 1998a:4. [see Blum et al. 1998b:428 ].
  Type - locality: Headwaters of the Gila, 6,000 ft.;  Emory, 18 Oct. 1846 (?). [Neotype: New Mexico, Grant Co., San Lorenzo;
    Benson 16660 , July 1966 (POM 317382) [Benson 1982:937].
  Nomenclatural synonyms:
    Mammillaria aggregata* Engelmann 1848b:157. nom. prov.
    Mammillaria aggregata* Engelmann ex Watson 1878:398. pro syn.
Echinocereus coccineus var. arizonicus (Orcutt) Ferguson 1989:221.
  = Echinocereus arizonicus.
Echinocereus coccineus var. arizonicus fa. matudae* Parker 1996:4 nom. nud.
  = Echinocereus matudae.
Echinocereus coccineus subsp. canyonensis* Blum 1997 Seed List nom. nud.
  = Echinocereus canyonensis.
Echinocereus coccineus var. conoideus* Weniger 1970:42 nom. inval. (no basionym given).
  = Echinocereus conoideus.
Cereus coccineus var. cylindricus Engelmann 1849:51.
  Type - locality: New Mexico, Mtns. near Santa Fe, Fendler; 17 Sept. 1846 (MO).
    Note: This plant is usually considered as a synonym of Echinocereus coccineus subsp. coccineus and has never been recombined
      under the genus Echinocereus.
Echinocereus coccineus var. gurneyi (Benson) Heil & Brack 1988:26.
  = Echinocereus triglochidiatus var. gurneyi.
Echinocereus coccineus subsp. gurneyi* Blum 1997 Seed List nom. nud.
  = Echinocereus triglochidiatus var. gurneyi .
Echinocereus coccineus subsp. gurneyi (L. Benson) D. Felix & H. Bauer 2014. - EcJ-Online 2 Sonderheft: XIII
  = Echinocereus triglochidiatus var. gurneyi .
Echinocereus coccineus fa. hexaedrus* Parker 1996:4 nom. nud.
  = Echinocereus hexaedrus.
Echinocereus coccineus var. inermis (Schumann) Purpus 1925:49.
  basionym: Echinocereus phoeniceus var. inermis Schumann 1896a:150.
    Type - locality: Colorado, Mesa Grande, Tongue Creek; C. A. Purpus (?).
      Nomenclatural synonyms:
        Echinocereus triglochidiatus var. inermis (Schumann) Arp 1973:132-133.
        Echinocereus triglochidiatus var. mojavensis fa. inermis (Schumann) Ferguson 1989:219.
        Echinocereus triglochidiatus var. melanacanthus fa. inermis* Alabaster 1994:105 nom. inval.
        Echinocereus coccineus var. mojavensis fa. inermis* Parker 1996:4 nom. nud.
        Echinocereus mojavensis fa. inermis* Mesa Garden Seed Catalog 1996 nom. nud.
        Echinocereus triglochidiatus subsp. mojavensis fa. inermis Blum in Blum et al. 1998a:9. [see Blum et al. 1998b:365].
Echinocereus coccineus var. kunzei (Gurke) Backeberg 1960:2070.
  = Echinocereus kunzei.
Cereus coccineus var. melanacanthus Engelmann 1849:51.
  Type - locality: New Mexico, Santa Fe; Fendler, 26 Nov. 1846 (MO).
    Nomenclatural synonyms:
      Echinocereus triglochidiatus var. melanacanthus (Engelmann) Benson 1944:254.
        Note: This is the name Benson applies to Echinocereus coccineus.
Echinocereus coccineus fa. lloydii* Parker 1996:4 nom. nud.
  = Echinocereus lloydii.
Echinocereus coccineus var. mojavensis* Parker 1996:4 nom. nud.
  = Echinocereus mojavensis.
Echinocereus coccineus subsp. mojavensis (Engelmann & Bigelow) Taylor 1997:8.
  = Echinocereus mojavensis.
Echinocereus coccineus var. mojavensis fa. inermis* Parker 1996:4 nom. nud.
  = Echinocereus coccineus var. inermis.
Echinocereus coccineus var. neomexicanus* Parker 1996:4 nom. nud.
  = Echinocereus neomexicanus.
Echinocereus coccineus var. neomexicanus fa. rosei* Parker 1996:4 nom. nud.
  = Echinocereus rosei.
Echinocereus coccineus var. nudum* Auct ? (CSJ 35:194) nom. nud.
Echinocereus coccineus var. octacanthus* Boissevain & Davidson 1940:39 nom. illegit.
  = Echinocereus octacantus (Muehlenpfordt) Britton & Rose 1922. [based on Echinopsis octacanthus Muehlenpfordt January,
    1848].
      Note: Echinocereus coccineus Engelmann 1848 was published in Wislizenus April 2, 1848.
Echinocereus coccineus var. paucispinus (Engelmann) Ferguson 1989:222.
  = Echinocereus paucispinus.
Echinocereus coccineus subsp. paucispinus Blum, Lange & Rutow in Blum et al. 1998a:4. [see Blum et al. 1998b:433].
  = Echinocereus paucispinus.
Echinocereus coccineus fa. roemeri* Parker 1996:4 nom. nud.
  = Echinocereus roemeri.
Echinocereus coccineus subsp. roemeri Blum, Lange & Rutow in Blum et al. 1998a:4. [see Blum et al. 1998b:426].
  = Echinocereus roemeri.
Echinocereus coccineus fa. rosei* Blum 1990:13. nom. inval. (no basionym given).
  = Echinocereus rosei.
Echinocereus coccineus subsp. rosei Blum & Rutow in Blum et al. 1998a:4. [see Blum et al. 1998b:426].
  = Echinocereus rosei.
Echinocereus coccineus var. rosei Zimmerman ex Powell & Weedin 2004:203.
  = Echinocereus rosei.
Echinocereus coccineus subsp. santaritensis (W. Blum & Rutow) M.A.Baker in Felger et al.2014:38.
  = Echinocereus santaritensis
Echinocereus coccineus subsp. transpecosensis Blum & Oldach 2015:52-64.
  Type - locality: Texas, Hudspeth Co., North of Sierra Blanca; Powell 6240 (SRSC)
Cereus concolor Schott1857:ii .
  Type - locality: Texas, Pecos County, Escondido Springs; Schott (?).
    Note: Apparently a form of Echinocereus chloranthus that has never been formally transfered to the genus Echinocereus.
Echinocereus congestispinus* Engelmann ex Haage 1859:19 nom. nud.
Echinocereus congestispinus* Forster ex Seitz 1870:10 nom. nud.
Echinocereus conglomeratus* Forster ex Seitz 1870:10 nom. nud.
Echinocereus conglomeratus Blanc 1890:56.
  Type - locality: Nuevo Leon, Rinconada; Mathsson, 24 Sept. 1889 (?) [according to Schumann 1897:278].
    Nomenclatural synonyms:
      Echinocereus bolansis* Runge ex Haage 1892:117 pro syn.
      Cereus conglomeratus (Blanc) Berger 1905:81.
      Echinocereus enneacanthus var. conglomeratus (Blanc) Benson 1974:80.
      Echinocereus stramineus var. conglomeratus (Blanc) Bravo1974:47.
      Echinocereus stramineus fa. conglomeratus* Parker 1996:3 nom. nud.
        Note: This is usually listed as Echinocereus conglomeratus Forster ex Schumann 1897:278. It is usually considered a
          taxonomic synonym of Echinocereus stamineus subsp. stramineus
Echinocereus conglomeratus var. robustior* Haage 1892:117 nom. nud.
Echinocereus conoideus (Engelmann & Bigelow) Engelmann ex Rumpler 1886:807.
  basionym: Cereus conoideus Engelmann & Bigelow in Engelmann 1856:284. [see Engelmann 1857:35].
    Type - locality: New Mexico, Anton Chico; J. M. Bigelow, 24 Sept. 1853 (MO).
      Nomenclatural synonyms:
        Cereus phoeniceus subsp. conoideus (Engelmann & Bigelow) Engelmann 1857:37.
        Echinocereus phoeniceus* var. conoideus Schumann 1896:150.
Echinocereus coccineus var. conoideus* Weniger 1970:42 nom. inval.
  Note: This species is usually treated as a taxonomic synonym of Echinocereus coccineus.
Echinocereus cowperi* Hort ex Blum et al. 1998b:208 pro syn. Echinocereus chloranthus subsp. rhyolithensis.
Echinocereus crispatus* Mathsson 1890:464 nom. nud.

Echinocereus ctenoides – Echinocereus monacanthus

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Echinocereus ctenoides* Lemaire 1868:57 nom. nud.
  [Probably = Echinocereus ctenoides Engelmann ex Rumpler].
Echinocereus ctenoides (Engelmann) Engelmann ex Rumpler 1886:819.
  basionym: Cereus ctenoides Engelmann 1856:279. [see Engelmann 1859:31].
    Type - locality: Coahuila, between Eagle Pass & Santa Rosa; Bigelow, Jan. 1853 (MO).
      Nomenclatural synonyms:
        Echinocereus dasyacanthus var. ctenoides (Engelmann) Backeberg1960:2021.
        Echinocereus pectinatus var. ctenoides* Weniger 1971:31 nom. inval. (no basionym given).
        Echinocereus pectinatus var. ctenoides (Engelmann) Weniger ex Frank 1991:135.
        Echinocereus pectinatus subsp. ctenoides (Engelmann) Frank 1997:136.
        Echinocereus spinosissimus var. ctenoides* (Engelmann) Ito 1981:237. nom. inval. (no basionym).
          Note: Echinocereus pectinatus var. wenigeri is considered by Frank (1997) to be a taxonomic synonym of this species.
Echinocereus cucumis Werdermann 1949:3-4.
  Type - locality: unknown [received from Hummel in 1937 without location noted, but probably from Sonora, Mexico].
    Nomenclatural synonyms:
      Echinocereus noctiflorus* Hort. ex Backeberg 1960:1991 pro syn.
      Echinocereus nocturniflorus* Hort. ex Backeberg 1960:1991 pro syn.
      Echinocereus scheeri var. gentryi cultivar 'Cucumis' (Werdermann) Taylor 1984:155.
      Echinocereus scheeri var. gentryi fa. cucumis* Parker 1996:4 nom. nud.
        Note: Usually considered a taxonomic synonym of Echinocereus scheeri subsp. gentryi.
Echinocereus dahliaflorus*.
  [An April fool prank name in Mollers Deutche Garter-Zeitung 15:148 (1900) ].
Echinocereus dasyacanthus Engelmann 1848a:100.
  Type - locality: Mexico, Chihuahua, Mtns. near Cd. Juarez; Wislizenus (MO).
    Nomenclatural synonyms:
      Cereus dasyacanthus (Engelmann) Engelmann 1849:50.
      Cereus deflexispinus* Monville ex Labouret 1853:321 pro syn.
      Echinocereus pectinatus var. dasyacanthus (Engelmann) Engelmann ex Haage 1892:118.
      Echinocereus pectinatus var. dasyacanthus* Earle 1971:80 nom. illegit. (later homonym).
      Echinocereus pectinatus var. dasyacanthus* Taylor 1984:179 nom. illegit. (later homonym).
        Note: Benson uses the name Echinocereus pectinatus var. neomexicanus for this species. Neotype: Wright (MO 2016859)
Echinocereus dasyacanthus subsp. crockettianus D. Felix & H. Bauer 2014. EcJ-Online 2 Sonderheft: V
  Type - locality: Texas: Crockett County, near Fort Lancaster; Anonymous 000577; 1991 (JE42/14)
Echinocereus dasyacanthus var. ctenoides (Engelmann) Backeberg 1960:2021.
  = Echinocereus ctenoides.
Echinocereus dasyacanthus ssp. ctenoides (Engelmann) Lode, Cact.-Avent. Int. 100:29, 2013.
  = Echinoereus ctenoides.
Echinocereus dasyacanthus var. hildmannii* Arendt ex Weniger 1970:33 nom. inval. (no basionym given)
  = Echinocereus hildmannii.
Cereus dasyacanthus var. minor* Engelmann 1856:278 nom. illegit.
  = Echinocereus roetteri.
    Note: This name is rejected by the author in his addenda of corrections and replaced by Echinocereus roetteri.
Echinocereus dasyacanthus subsp. multispinosus D. Felix & H. Bauer 2014. - EcJ-Online 2 Sonderheft: VII
  Type - locality: Mexico: Chihuahua; west of  Sierra El Morrion; Anonymous 002236; 1997 (JE 41/14)
Cereus dasyacanthus var. neomexicanus Coulter 1896:384.
  Type - locality: "stony prairies along Escondido Creek", [Pecos County, Texas]; Wright 366, 6 June 1851 (MO).
    Nomenclatural synonyms:
      Echinocereus pectinatus var. neomexicanus (Coulter) Benson 1944:256.
      Echinocereus dasyacanthus var. neomexicanus* Coulter ex Weniger 1970:32 nom. inval.
        Note: Benson uses the name Echinocereus pectinatus var. neomexicanus for Echinocereus dasyacanthus subsp.
          dasyacanthus. The original var. neomexicanus was probably a natural hybrid (see Zimmerman 1993).
Echinocereus dasyacanthus var. neomexicanus* Coulter ex Weniger 1970:32 nom. inval. (no basionym given).
Echinocereus dasyacanthus var. rectispinus Trocha & Fethke 1991:31-40.
  Type - locality: Mexico, Chihuahua, near Villa Ahumada; Fiehn 1979. (Hamburg Herbarium).
    Nomenclatural synonyms:
      Echinocereus pectinatus var. rectispinus* Zimmerman 1993:277 pro syn.
      Echinocereus dasyacanthus subsp. rectispinus (Trocha & Fethke) Blum, Rischer & Rutow in Blum .
Echinocereus dasyacanthus subsp. rectispinus (Trocha & Fethke) Blum, Rischer & Rutow in Blum et al. 1998a:2. [see Blum et al. 1998b:118].
  = Echinocereus dasyacanthus var. rectispinus Trocha & Fethke 1991.
Echinocereus dasyacanthus var. rigidissimus (Engelmann) Marshall & Bock 1941:119.
  = Echinocereus pectinatus var. rigidissimus.
Echinocereus dasyacanthus fa. roetteri* Parker 1996:2 nom. nud.
  = Echinocereus roetteri.
Echinocereus dasyacanthus var. rubrus* Frič 1924:121 nom. nud.
Cereus dasyacanthus var. spurius Labouret 1853:321.
  Type - locality: unknown; ? (?).
    Nomenclatural synonyms:
      Cereus deflexispinus var. spurius Monville ex Labouret 1853:321. pro syn.
Echinocereus dasyacanthus var. steereae (Clover) Marshall 1945:115.
  = Echinocereus steereae.
Echinocereus dasyacanthus fa. steereae* Parker 1996:2 nom. nud.
  = Echinocereus steereae.
Echinocereus datae Laferriere 1997:28.
  Type - locality:Mexico, Chihuahua, near Nabogame; Laferriere 1425, 8 Apr 1988 (ARIZ).
Echinocereus davisii Houghton 1931:466.
  Type - locality: Texas, 4 mi. south of Marathon; 4000 ft., Houghton 700 (US 1566585).
    Nomenclatural synonyms:
      Echinocereus viridiflorus var. davisii (Houghton) Marshall & Bock 1941:119.
      Echinocereus viridiflorus fa. davisii (Houghton) Krainz 1967:51.
      Echinocereus viridiflorus subsp. davisii (Houghton) Taylor 1997:10.
Echinocereus decumbens Clover & Jotter 1941:417.
  Type - locality: Arizona, Coconino Co., Mile 16. ; Marble Canyon; Clover & Jotter 2212, 15 July 1938 (MICH 1115972).
    Nomenclatural synonyms:
      Echinocereus engelmannii var. decumbens (Clover & Jotter) Benson 1944:258.
      Echinocereus engelmannii subsp. decumbens (Clover & Jotter) Blum & Lange in Blum et al. 1998a:2.
Cereus deflexispinus* Monville ex Labouret 1853:321 pro syn.
  = Echinocereus dasyacanthus.
Cereus deflexispinus* var. spurius Monville ex Labouret 1853:321 pro syn.
  = Cereus dasyacanthus var. spurius.
Echinocereus degandii* Rebut ex Schumann 1895:123 nom. nud.
  [Probably = Echinocereus dasyacanthus].
Echinocereus delaetii (Gurke) Gurke 1909b:131.
  basionym: Cephalocereus delaetii Gurke 1909a:116-121.
    Type - locality: unknown [Probably Sierra Paila, Coahuila].
      Nomenclatural synonyms:
        Echinocereus longisetus var. delaetii (Gurke) Taylor 1988:79.
        Echinocereus longisetus subsp. delaetii (Gurke) Taylor 1997:9.
Echinocereus delaetii var. freudenbergeri (Frank) Taylor 1985b:97.
  = Echinocereus freudenbergeri.  [see Echinocereus longisetus subsp. freudenbergii ].
Cereus deppei Salm-Dyck 1834:338.
  Type - locality: Mexico; ? (?).
    Note: This species is usually considered a taxonomic synonym of Echinocereus cinerascens.
Cereus deppei* Hort. Berol. ex Pfeiffer 1837:101 pro syn.
  = Echinocereus cinerascens.
Cereus deppei* Hort. Paris ex Salm-Dyck 1845:27 nom. nud.
Echinocereus deppei *(Salm-Dyck) Schumann 1895:123 pro syn.
  = Echinocereus cinerascens.
Echinocereus depressus* Haage 1900:157 nom. nud.
*Echinocereus diguetii* Hort. nom. nud. = Peniocereus diguetii.
Echinocereus dubius (Engelmann) Engelmann ex Rumpler 1886:787.
  basionym: Cereus dubius Engelmann 1856:282-283. [see in Engelmann 1859:36].
    Type - locality: Texas, Hudspeth Co. near Rio Grande; Wright 410, 19 June 1852 (MO 3347359).
      Nomenclatural synonyms:
        Echinocereus malibrani* Rebut ex Haage 1892:118.
        Echinocereus enneacanthus var. dubius (Engelmann) Benson 1969d:127.
        Echinocereus enneacanthus fa. dubius* Parker 1996:2 nom. nud.
          Note: This species is usually considered to be a taxonomic synonym of Echinocereus enneacanthus subsp. enneacanthus.
*Echinocereus dumenilianus Monville ex Haage 1859:19 = Echinopsis sp.
Echinocereus dumesnilianus* Walton 1898:26  pro syn. = Echinocereus candicans var. dumesnilianus*
Echinocereus durangensis Poselger ex Rumpler 1886:799.
  Type - locality: Mexico, Durango; ? (?).
    Nomenclatural synonyms:
      Echinocereus acifer var. durangensis (Poselger ex Rumpler) Schumann 1897:287.
      Echinocereus polyacanthus var. durangensis * Parker 1996:4 nom. nud.
        Note: Taylor (1985b) lists this name as a synonym of Echinocereus polyacanthus.
Echinocereus durangensis var. nigrispinus Hort. ex Rumpler 1886:800.
  Type - locality: Mexico, Durango; ? (?).
Echinocereus durangensis var. rufispinus Hort. ex Rumpler 1886:800.
  Type - locality: Mexico, Durango; ? (?).
Echinocereus ehrenbergii* Engelmann ex Haage 1859:19. nom. nud.
  [Probably = Echinocereus ehrenbergii (Pfeiffer) Pfeiffer ex Rumpler].
Echinocereus ehrenbergii (Pfeiffer) Pfeiffer ex Rumpler 1886:775.
  basionym: Cereus ehrenbergii Pfeiffer 1840:282.
    Type - locality: Mexico, Hidalgo, Mineral del Monte [according to Schumann 1897:262]; Ehrenberg (?).
      Nomenclatural synonyms:
        Echinocereus pentalophus var. ehrenbergii (Pfeiffer) Backeberg 1960:2003.
        Echinocereus cinerascens var. ehrenbergii (Pfeiffer) Bravo 1974:47.
        Echinocereus cinerascens subsp. ehrenbergii (Pfeiffer) Blum & Rutow in Blum et al. 1998a:4.
*Echinocereus emoryi = Bergerocactus emoryi.
Echinocereus engelmannii* Lemaire 1868:56 nom. nud.
  [Probably = Echinocereus engelmannii Parry ex Rumpler].
Echinocereus engelmannii* Harry ex Seitz 1878:10. nom. nud.
  [Probably = Echinocereus engelmannii Parry ex Rumpler].
Echinocereus engelmannii (Parry ex Engelmann) Parry ex Rumpler 1886:805.
  basionym: Cereus engelmannii Parry ex Engelmann 1852:328.
    Type - locality: California, Mountains near San Felipe; Parry (?). [Neotype: California, San Felipe Valley; Benson 16386 (POM).
Echinocereus engelmannii var. acicularis Benson 1969a:22.
  Type - locality: Arizona Maricopa Co., Crossing of New River, south side of Black Canyon Refuge, 1300 ft.; Benson 16,616,
    20 Apr. 1966 (POM 311313).
Echinocereus engelmannii var. albispinus Carp 1894:29.
  Type - locality: unknown; ? (?).
Echinocereus engelmannii var. albispinus Cels ex Schumann 1897:276 nom. nud.
Echinocereus engelmannii var. armatus Benson 1969c:33.
  Type - locality: Calif., San Bernardino Co., Dead Man Point; Benson 14767, 1 Apr. 1951 (POM 284927).
Echinocereus engelmannii var. bonkerae* Mesa Garden Seed Catalog 1996 nom. nud.
  = Echinocereus bonkerae.
Echinocereus engelmannii var. boyce-thompsonii* Mesa Garden Seed Catalog 1996 nom. nud.
  = Echinocereus boyce-thompsonii.
Echinocereus engelmannii var. caespitosus* Orcutt 1897:276 nom. nud.
Echinocereus engelmannii var. chrysocentrus (Engelmann & Bigelow) Engelmann ex Rumpler 1886:806.
  basionym: Cereus engelmannii var. chrysocentrus Engelmann & Bigelow in Engelmann 1856:283. [see Englemann 1857:35].
    Type - locality: Arizona, Bill Williams River; J. M. Bigelow, 13 Feb. 1854 (MO).
      Nomenclatural synonyms:
        Echinocereus chrysocentrus (Engelmann & Bigelow) Orcutt 1926:4.
        Echinocereus engelmannii fa. chrysocentrus (Engelmann & Bigelow) Schelle 1907:135
Echinocereus engelmannii fa. chrysocentrus (Engelmann & Bigelow) Schelle 1907:135.
  = Echinocereus engelmannii var. chrysocentrus.
Echinocereus engelmannii var. decumbens (Clover & Jotter) Benson 1944:258.
  = Echinocereus decumbens.
Echinocereus engelmannii subsp. decumbens (Clover & Jotter) Blum & Lange in Blum et al. 1998a:2. [see Blum et al. 1998b:76].
  = Echinocereus decumbens.
Echinocereus engelmannii var. fasciculatus* Mesa Garden Seed Catalog 1996 nom. nud.
  = Echinocereus fasciculatus.
Echinocereus engelmannii subsp. fasciculatus (Engelmann ex Watson) Blum & Lange in Blum et al. 1998a:2.
  = Echinocereus fasciculatus.
    Note: see Blum et al. 1998b:78.
Echinocereus engelmannii var. fulvispinus* Cels ex Schumann 1897:276 nom. nud.
Echinocereus engelmannii var. howei Benson 1974:80.
  Type - locality: Calif., San Bernardino Co., east of Goffs; D. F. Howe 4570 (POM 317,886).
Echinocereus engelmannii subsp. llanuraensis (Rutow) Felger 1999: 20. Proc. San Diego Soc. Nat. Hist., 35: 20.
  =Echinocereus nicholii subsp. llanuraensis Rutow.
Echinocereus engelmannii var. munzii (Parish) Pierce & Fosberg 1933:123.
  = Echinocereus munzii.
Echinocereus engelmannii subsp. munzii* Blum 1997 Seed List nom. nud.
  = Echinocereus munzii.
Echinocereus engelmannii fa. munzii* (Parish) Ruinaard, Succulenta 89(3): , Nom. inval. (Art. 33.4),
  = Echinocereus munzii.
Echinocereus engelmannii var. nicholii Benson 1944:258.
  Type - locality: Arizona, Pima Co., Silver Bell Mtns.; Benson 10720, 28 Mar. 1941 (Ariz 34989).
    Nomenclatural synonyms:
      Echinocereus nicholii (Benson) Parfitt 1987:157-158.
      Echinocereus engelmannii subsp. nicholii* (Benson) Breckwoldt 1993:8 nom. inval.
Echinocereus engelmannii subsp. nicholii* (Benson) Breckwoldt 1993:8 nom. inval. (no basionym given)
  = Echinocereus engelmannii var. nicholii
Echinocereus engelmannii var. pfersdorffii* Heyder ex Schumann 1897:276 nom. nud.
Echinocereus engelmannii var. purpureus Benson 1969d:126-127.
  Type - locality: Utah, Washington Co., north of St. George, 2900 ft.; Benson 13637, 5 May 1949 (POM 285578).
Echinocereus engelmannii var. robustior* Haage 1892:117 nom. nud.
Echinocereus engelmannii var. robustior* Hildmann ex Hirscht 1894:194 nom. nud.
Echinocereus engelmannii var. variegatus (Engelmann & Bigelow) Engelmann ex Rumpler 1886:806.
  basionym: Cereus engelmannii var. variegatus Engelmann & Bigelow in Engelmann 1856:283. [see Engelmann 1857:35].
    Type - locality: Arizona, Cactus Mountains at head of Bill Williams River; J.M. Bigelow, 1 Feb. 1854 (MO 2016811).
      Nomenclatural synonyms:
        Echinocereus engelmannii subsp. variegatus (Engelmann & Bigelow) E. Lutz 2000:97.
Echinocereus engelmannii fa. variegatus (Engelmann & Bigelow) Schelle 1907:135.
  = Echinocereus engelmannii var. variegatus.
Echinocereus engelmannii subsp. variegatus* Breckwoldt 1993 in Kakteen und andere Sukkulenten 44(8) nom. inval.(No basionym)
  = Echinocereus engelmannii var. variegatus.
Echinocereus engelmannii subsp. variegatus ( Engelm. & J.M.Bigelow ) E. Lutz. 2000:97.
  = Echinocereus engelmannii var. variegatus.
Echinocereus engelmannii var. versicolor* Hirscht 1894:194 nom. nud.
Echinocereus engelmannii var. versicolor Walton 1898:118,
  Type - locality: California (?).
Echinocereus enneacanthus Engelmann 1848a:111.
  Type - locality: Mexico, Chihuahua, near San Pablo; Wislizenus 244, 8 Apr. 1847 (MO 83797).
    Nomenclatural synonyms:
      Cereus enneacanthus (Engelmann) Engelmann 1849:50.
        Note: Echinocereus lowryi* Lowry 1936:20 nom. nud. and Echinocereus albiflorus* Lowry 1937:20 nom. nud. are probably
          forms of this species.
Echinocereus enneacanthus fa. brevispinus Moore 1967:93-94.
  Type - locality: Texas, Starr Co., 8 miles East of Rio Grande City; Clover, 31 Dec 1953 (MICH).
    Nomenclatural synonyms:
      Echinocereus enneacanthus var. brevispinus (Moore) Benson 1969d:127.
      Echinocereus enneacanthus subsp. brevispinus (Moore) Taylor 1997:8.
Echinocereus enneacanthus var. brevispinus (Moore) Benson 1969d:127.
  = Echinocereus. enneacanthus fa. brevispinus.
Echinocereus enneacanthus subsp. brevispinus (Moore) Taylor 1997:8.
  = Echinocereus enneacanthus fa. brevispinus.
Echinocereus enneacanthus var. brevispinus fa. carnosus* Parker 1996:2 nom. nud.
  = Echinocereus carnosus.
Echinocereus enneacanthus fa. carnosus (Rumpler) Schelle 1907:127.
  = Echinocereus carnosus.
Echinocereus enneacanthus var. carnosus* Quehl 1908:114 nom. nud.
  [Probably = Echinocereus carnosus].
Echinocereus enneacanthus fa. carnosus* is attributed incorrectly to Matthson ex Schumann by Bravo 1991:76.
Echinocereus enneacanthus var. conglomeratus (Blanc) Benson 1974:80.
  = Echinocereus conglomeratus.
Echinocereus enneacanthus var. dubius (Engelmann) Benson 1969d:127.
  = Echinocereus dubius.
Echinocereus enneacanthus fa. dubius* Parker 1996:2 nom. nud.
  = Echinocereus dubius.
Echinocereus enneacanthus var. erectus* Haage 1892:117 nom. nud.
Echinocereus enneacanthus fa. intermedius Moore 1967:93.
  Type - locality: Texas, Starr Co., off La Grulla Road, Clover, 23 Mar. 1953 (MICH 1133318).
    Note: Usually considered a taxonomic synonym of Echinocereus enneacanthus subsp. brevispinus.
Echinocereus enneacanthus fa. major* Schelle 1907:127 nom. nud.
Echinocereus enneacanthus fa. major* Hort. ex Borg 1951:223 nom. inval.
Echinocereus enneacanthus var. majus* Haage 1892:117 nom. nud.
Echinocereus enneacanthus fa. merkeri* Parker 1996:2 nom. nud.
  = Echinocereus merkeri.
Echinocereus enneacanthus fa. sarissophorus* Parker 1996:2 nom. nud.
  = Echinocereus sarissophorus.
Echinocereus enneacanthus var. stramineus (Engelmann) Benson 1969d:127.
  = Echinocereus stramineus.
Echinocereus enneacanthus fa. uspenskii* Parker 1996:2 nom. nud.
  = Echinocereus uspenskii.
*Echinocereus erectocentrus var. pallidus error in spelling = Echinomastus erectocentrus.
Echinocereus fasciculatus (Engelmann) Benson 1969a:21.
  basionym: Mammillaria fasciculata* Engelmann in Emory 1848:156. nom. prov.
    Type - locality: Arizona, Graham Co. 6-7 miles east of Solomon. [Neotype - Benson 16633, 22 April 1966 (POM 311339) ].
      Nomenclatural synonyms:
        Echinocereus fendleri var. fasciculatus (Engelmann) Taylor 1985a:252.
        Echinocereus engelmannii var. fasciculatus* Mesa Garden Seed Catalog 1996 nom. nud.
        Echinocereus engelmannii subsp. fasciculatus (Engelmann ex Watson) Blum & Lange in Blum et al. 1998a:2.
          Note: The validity of the name Echinocereus fasciculatus is uncertain. Echinocereus robustus is usually considered to be a
            taxomnomic synonym of this plant. The  name Mammillaria fasciculata is listed as a synonym of Cereus fendleri by
              Watson 1878:404.
Echinocereus fasciculatus subsp. abbeae* Blum 1997 Seed List nom. nud.
  = Echinocereus abbeae.
Echinocereus fasciculatus var. bonkerae (Thornber & Bonker) Benson 1969a:22.
  = Echinocereus bonkerae.
Echinocereus fasciculatus subsp. bonkerae (Thornber & Bonker) Taylor 1997:8.
  = Echinocereus bonkerae.
Echinocereus fasciculatus var. boyce-thompsonii (Orcutt) Benson 1969a:21.
  = Echinocereus boyce-thompsonii.
Echinocereus fasciculatus subsp. boyce-thompsonii (Orcutt) Taylor 1997:8.
  = Echinocereus boyce-thompsonii.
Echinocereus fasciculatus var. rectispinus* Benson 1969a:131 pro syn.
  = Echinocereus rectispinus.
Echinocereus fasciculatus fa. robustus* Parker 1996:2 nom. nud.
  = Echinocereus robustus.
Echinocereus felixianus H. Bauer 2014. - EcJ-Online 2 Sonderheft: IX
  Type - locality: Mexico: Coahuila: Municipio Cuatro Cienegas de Carranza; Anonymous 005387; 2010 (JE 42/14).
Echinocereus fendleri* Sencke ex Seitz 1870:11 nom. nud.
  [Probably a synonym of Echinocereus fendleri Loder].
Echinocereus fendleri (Engelmann) Loder 1882:445.
  basionym: Cereus fendleri Engelmann 1849:51.
    Type - locality: New Mexico, near Santa Fe; Fendler 273, June 1846 (MO 2016839).
      Nomenclatural synonyms:
        Echinocereus melaleuca* Foster ex Haage 1892:117 nom. nud.
        Cereus cinerascens var. fendleri (Engelmann) Bois 1928:371.
          Note: Usually listed as Echinocereus fendleri Engelmann ex Rumpler 1886:801 or Senke ex Haage. 1893:22.
Echinocereus fendleri var. albiflorus (Weingart) Backeberg 1960:2047.
  = Echinocereus albiflorus.
Echinocereus fendleri fa. albiflorus* Parker 1996:2 nom. nud.
  = Echinocereus albiflorus Weingart.
Echinocereus fendleri var. bonkerae (Thornber & Bonker) Benson 1944:260
  = Echinocereus bonkerae.
Echinocereus fendleri var. boyce-thompsonii (Orcutt) Benson 1944:260.
  = Echinocereus boyce-thompsonii.
Echinocereus fendleri var. fasciculatus (Engelmann) Taylor 1985a:252.
  = Echinocereus fasciculatus.
Echinocereus fendleri var. hempelii* Mesa Garden Seed Catalog 1996 nom. nud.
  = Echinocereus hempelii.
Echinocereus fendleri subsp. hempelii Blum in Blum et al. 1998a:2. [see Blum et al. 1998b:93].
  = Echinocereus hempelii.
Echinocereus fendleri var. kuenzleri (Castetter et al.) Benson 1982:942.
  = Echinocereus kuenzleri.
Echinocereus fendleri subsp. kuenzleri* Blum 1997 Seed List nom. nud.
  = Echinocereus kuenzleri.
Echinocereus fendleri subsp. kuenzleri (Castetter et al.) W. Blum & D. Felix. 2014 - EcJ-Online 2 (2): 105.
  = Echinocereus kuenzleri.
Echinocereus fendleri var. ledingii (Peebles) Taylor 1985a:253.
  = Echinocereus ledingii.
Echinocereus fendleri var. major* Haage 1900:158 nom. nud.
Echinocereus fendleri var. major Rother 1930:109.
  Type - locality: Mexico (?)
Echinocereus fendleri var. pauperculus (Engelmann) Schumann 1897:275.
  basionym: Cereus fendleri var. pauperculus Engelmann 1849:51.
    Type - locality: New Mexico, near Santa Fe; Fendler, 4 Nov. 1846 (MO).
Echinocereus fendleri var. rectispinus (Peebles) Benson 1944:259.
  = Echinocereus rectispinus.
Echinocereus fendleri subsp. rectispinus (Peebles) Taylor 1997:9.
  = Echinocereus rectispinus.
Echinocereus fendleri var. robustus Fobe 1911:55 [not Benson 1944].
  Type - locality: unknown; ? (?).
Echinocereus fendleri var. robustus* (Peebles) Benson 1944:259 nom. illegit. (later homonym).
  = Echinocereus robustus.
Echinocereus fereirae*  Gates 1935:154 nom. prov. for Echinocereus ferreirianus Gates 1953. [ = Gates 550]
Echinocereus ferreirianus Gates 1953:8-11.
  Type - locality: Mexico, Baja Calif. Norte, Isla Piojo, Bahia de los Angeles; H. E. Gates, 1934 (DS 375348).
Echinocereus ferreirianus var. lindsayi (Meyran) Taylor 1985b:46.
  = Echinocereus lindsayi.
Echinocereus ferreirianus subsp. lindsayi (Meyran) Taylor 1997:9.
  = Echinocereus lindsayi.
Echinocereus ferreirrae* Gates 1935:154 nom. nud. [= Gates number 550].
  = Echinocereus ferreirianus.
Echinocereus fiehnii* Lau 1992:74 nom. nud.
  = Echinocereus russanthus var. fiehnii.
Echinocereus finnii* Hort. ex Taylor 1985 101 pro syn.
  = Echinocereus russanthus var. fiehnii.
Echinocereus fitchii Britton & Rose 1922:30.
  Type - locality: Texas, near Laredo; Rose 18037, 1913 (US 1821071).
    Nomenclatural synonyms:
      Echinocereus reichenbachii var. fitchii (Britton & Rose) Benson 1969d:127.
      Echinocereus reichenbachii subsp. fitchii (Britton & Rose) Taylor 1997:9.
Echinocereus fitchii var. albertii* Mesa Garden Seed Catalog 1996 nom. nud.
  = Echinocereus reichenbachii var. albertii.
Echinocereus fitchii subsp. albertii (Benson) Blum & Lange in Blum et al. 1998a:5. [see Blum et al. 1998b:313].
  = Echinocereus reichenbachii var. albertii.
Echinocereus fitchii var. armatus* Mesa Garden Seed Catalog 1996 nom. nud.
  = Echinocereus pectinatus var. armatus.
Echinocereus fitchii subsp. armatus (Poselger) Blum, Lange & Rutow in Blum et al. 1998a:5. [see Blum et al. 1998b:314].
  = Echinocereus pectinatus var. armatus.
Echinocereus fitchii subsp. bergmannii Blum 2005:265.
  Type - locality: Mexico, Tamaulipas, Sierra de San Carlos; Bergmann 221, 5 Sept. 2003 (UAT 021650).
*Echinocereus flavescens = Haageocereus sp.
Echinocereus flavidiflorus* Haage 1892:117 nom. nud.
Echinocereus flavidiflorus* Hildmann ex Schumann 1897:264 pro syn.
  = Echinocereus leonensis.
Echinocereus flaviflorus (Coulter) Coulter ex Schumann 1897:274.
  basionym: Cereus flavidiflorus Engelmann ex Coulter 1896:391.
    Type - locality: Mexico, Baja California del Norte, west of San Borgia; Gabb 10, 1867 (MO).
      Note: Usually considered a synonym of Echinocereus maritimus.
Echinocereus flavispinus* Bailey 1941:270 nom. nud.
Echinocereus floresii Backeberg 1949:5.
  Type - locality: Mexico, Sinaloa, near Topolobampo; R. Flores & F. Schwarz (?).
    Nomenclatural synonyms:
      Echinocereus sciurus var. floresii (Backeberg) Taylor1985b:115.
      Echinocereus sciurus subsp. floresii (Backeberg) Taylor1997:10.
      Echinocereus bristolii subsp. floresii Blum & Lange in Blum et al. 1998a:3. [see Blum et al. 1998b:353].
Echinocereus fobeanus Oehme 1940:49-50.
  Type - locality: unknown [Probably Coahuila]; ? (?).
    Nomenclatural synonyms:
      Echinocereus chisoensis var. fobeanus (Oehme) Taylor 1985a:261.
      Echinocereus chisoensis subsp. fobeanus (Oehme) N.P. Taylor 2005 Cactaceae Syst. Init. 19: 10.
        Note: Taylor (1993) lists Echinocereus metornii as a taxonomic synonym of this species.
Echinocereus fobeanus subsp. metornii (Frank) Blum & Lange in Blum et al. 1998a:5. [see Blum et al. 1998b:307.]
  = Echinocereus metornii.
Echinocereus fordtii* spelling error for Echinocactus fordtii.
Echinocereus forsteri* Haage 1859:19. nom. nud.
Echinocereus freudenbergeri Frank 1981:102-105.
  Type - locality: Mexico, Coahuila, 12 km south of Cuatro Cienegas; G. Freudenberger (ZSS AA18-27).
    Nomenclatural synonyms:
      Echinocereus delaetii var. freudenbergeri (Frank) Taylor 1985b:97.
      Echinocereus setosus* Hort. ex Taylor 1988:76 pro syn.
      Echinocereus nivosus var. freudenbergeri* Parker 1996:3 nom. nud.
      Echinocereus longisetus subsp. freudenbergeri (Frank) Blum in Blum et al. 1998a:6. [see Blum et al. 1998b:187 ].
        Note: Taylor (1993) lists this species as a taxonomic synonym of Echinocereus nivosus.
Echinocereus galtieri* Hort. Grus. ex Schumann 1895:124 nom. nud.
Echinocereus galtieri Rebut ex Fobe 1911:56.
  Type - locality: unknown; ? (?).
    Nomenclatural synonyms:
      Echinocereus polyacanthus var. galtieri (Rebut ex Fobe) Schelle 1926:179.
Echinocereus gentryi Clover 1938:565.
  Type - locality: Mexico, Sonora, Canon Saucito; H.S. Gentry, 1932 (MICH).
    Nomenclatural synonyms:
      Echinocereus scheeri var. gentryi (Clover) Taylor 1984:154.
      Echinocereus scheeri subsp. gentryi (Clover) Taylor 1997:9
        Note: Echinocereus cucumis is usually considered a taxonomic synonym of this species.
*Echinocereus gladiatus = Echinopsis gladiata.
Echinocereus glycerius* Forster ex Haage 1859:19 nom. nud.
Echinocereus glycimorphus* Forster ex Seitz 1870:11 nom. nud.
Echinocereus glycimorphus Forster ex Rumpler 1886:800.
  Type - locality: unknown [Hidalgo near Ixmiquilpan and El Cardonal according to Schumann 1897:262] ; ? (?).
    Nomenclatural synonyms:
      Cereus glycimorphus (Forster ex Rumpler) Orcutt 1903:5.
      Echinocereus cinerascens fa. glycimorphus* Parker 1996:2 nom. nud.
Echinocereus glycimorphus var. undulatus Schelle 1926:166.
  Type - locality: unknown; ? (?).
    Note: Schelle 1926:166 lists Echinocereus undulatus* Hildmann as a synonym of this variety.
      Cereus glomeratus* Orcutt 1897:21 pro syn.= Echinocereus maritimus.
Echinocereus gonacanthus* Lemaire 1868:57 nom. nud.
  [Probably = Echinocereus gonacanthus Loder].
Echinocereus gonacanthus (Engelmann & Bigelow) Loder 1882:444.
  basionym: Cereus gonacanthus Engelmann & Bigelow in Engelmann 1856:283. [see Engelmann 1857:33].
    Type - locality: Arizona, Apache Co., Jacobs Well, 40 mi. west of Zuni; J. M. Bigelow, 29 Nov. 1853 (MO).
      Nomenclatural synonyms:
        Echinocereus paucispinus var. gonacanthus (Engelmann & Bigelow) Schumann 1897:281.
        Echinocereus triglochidiatus var. gonacanthus (Engelmann & Bigelow) Boissevain 1940:36.
          Note: Usually listed as Echinocereus gonacanthus Engelmann ex Rumpler 1886:806.
*Echinocereus grahamii (Monatschrift Kakteenkunde 20:47) error for Mammillaria grahamii.
Echinocereus grandis Britton & Rose 1922:18.
  Type - locality: Mexico, Gulf of California, San Esteban Island; J.N. Rose 16823, 13 Apr. 1911 (US).
    Nomenclatural synonyms:
      Echinocereus pectinatus var. grandis (Britton & Rose) Marshall 1955:68.
Echinocereus hancockii Dawson 1949:89-93.
  Type - locality: Mexico, Baja California Sur, north side of San Hipolito Bay; E. Y. Dawson 6443, 4 April 1949 (LAM).
    Nomenclatural synonyms:
      Echinocereus maritimus var. hancockii (Dawson) Taylor 1985b:44.
      Echinocereus maritimus subsp. hancockii (Dawson) Blum & Rutow in Blum et al. 1998a:6 [see Blum et al. 1998b:57 ].
Echinocereus havermansii* Rebut ex Haage 1892:118 nom. nud.
Echinocereus havernicki* Hildm. ex Schwarzbach & Bitter 1915:8 nom. nud.
Echinocereus hempelii Fobe 1897:187.
  Type - locality: Mexico [Probably Sierra del Nido, Chihuahua, Mexico] ; ? (?).
    Nomenclatural synonyms:
      Echinocereus fendleri var. hempelii * Mesa Garden Seed Catalog 1996 nom. nud.
      Echinocereus fendleri subsp. hempelii Blum in Blum et al. 1998a:2.
        Note: Some authors treat Echinocereus kuenzleri as a taxonomic synonym of this species.
Echinocereus hempelii var. centrispinus* Hort. nom. nud.
  [=HK 1489 from San Buenaventura, Chihuahua, Mexico].
Echinocereus hexaedrus (Engelmann & Bigelow) Engelmann ex Rumpler 1886:807.
  basionym: Cereus hexaedrus Engelmann & Bigelow in Engelmann 1856:285. [see Engelmann 1857:34].
    Type - locality: Arizona, Apache County, 15 miles west of Zuni; J. M. Bigelow, 28 Nov. 1853 (MO).
      Nomenclatural synonyms:
        Echinocereus paucispinus var. hexaedrus (Engelmann & Bigelow) Schumann 1897:281.
        Echinocereus triglochidiatus var. hexaedrus (Engelmann & Bigelow) Boissevain 1940:36.
        Echinocereus coccineus fa. hexaedrus* Parker 1996:4 nom. nud.
          Note: Considered by Taylor (1985b) as a taxonomic synonym of Echinocereus triglochidiatus var. melanacanthus.
Echinocereus hidalgensis* Hort. Berol. ex Backeberg 1962:3855-3856 nom. nud.
Echinocereus hildmannii Arendt 1892:146.
  Type - locality: unknown [West Texas according to Schelle 1926:175] ; ? (?).
    Nomenclatural synonyms:
      Echinocereus dasyacanthus var. hildmannii* Arendt ex Weniger 1970:33 nom. inval.
        Note: This plant may be a natural hybrid (see Zimmermann 1993).
Echinocereus huitcholensis (Weber) Gurke 1906:23.
  basionym: Cereus huitcholensis Weber 1904:383.
    Type - locality: Mexico, Jalisco, Sierra de los Huitcholes, 1800 m; L. Diguet, May 1900 (P).
      Nomenclatural synonyms:
        Echinocereus polyacanthus var. huitcholensis (Weber) Taylor1988:82.
        Echinocereus acifer subsp. huitcholensis (Weber) Lange 1994:109.
        Echinocereus polyacanthus var. acifer fa. huitcholensis*Parker 1996:4 nom. nud.
        Echinocereus polyacanthus subsp. huitcholensis (Weber) Taylor 1997:9.
*Echinocereus hypogaeus = Eridisia spiniflora.
Echinocereus inermis* Haage ex Hirscht 1898:130 pro syn.
  = Echinocereus knippelianus.
*Echinocereus intricatus = Echinopsis strigosa.
Echinocereus jacobyi* Hort. ex Schumann 1897:278 pro syn.
  = Echinocereus merkeri.
Echinocereus joconosle* Haage 1892:117 nom. nud.
Echinocereus klapperi Blum in Blum et al. 1998a:5. [see Blum et al. 1998b:380-383].
  Type - locality:Mexico, Sonora, El Novillo,500 m., Klapper 0121, 12 March 1983 (ZSS A 17985).
    Nomenclatural synonyms:
      Echinocereus klapperi subsp. bacanorensis (Rischer & Trocha) Rischer 2003:22.
Echinocereus klapperi subsp. bacanorensis (Rischer & Trocha) Rischer 2003:22.
  = Echinocereus salm-dyckianus subsp. bacanorensis Rischer & Trocha.
Echinocereus knippelianus Liebner 1895:170.
  Type - locality: unknown [Relocated in Sierra Parras, Coahuila, Mexico] ; ? (?).
    Nomenclatural synonyms:
      Echinocereus liebnerianus* Liebner ex Schumann 1896b:262 nom. nud.
      Echinocereus inermis* Haage ex Hirscht 1898:130 pro syn.
      Cereus knippelianus (Liebner) Orcutt 1902:27.
Echinocereus knippelianus subsp. kaplanii Halda & Stadkovshy 2000:34.
  Type - locality: Mexico, Nuevo Leon, North West of Los Mimbres alt. 2510 m.; Halda & Stadkovshy 8504132 (PR).
Echinocereus knippelianus var. kruegeri Glass & Foster 1978b:80.
  Type - locality: Mexico, Nuevo Leon, north of Ascencion near San Juanito; Glass & Foster 3902, 1 Mar. 1972 (POM).
Echinocereus knippelianus subsp. kruegeri (Glass & Foster) Blum & Lange in Blum et al. 1998b:248.
  = Echinocereus knippelianus var. kruegeri.
Echinocereus knippelianus var. kruegeri fa. reyesii* Parker 1996:3 nom. nud.
  = Echinocereus knippelianus var. reyesii.
Echinocereus knippelianus var. reyesii Lau 1980:264-265.
  Type - locality: Mexico, Nuevo Leon, between Siberia & Encantada; Lau 1237A, Apr. 1977 (POM).
    Nomenclatural synonyms:
      Echinocereus knippelianus var. kruegeri fa. reyesii*Parker 1996:3 nom. nud.
Echinocereus knippelianus subsp. reyesii (Lau) Blum & Lange in Blum et al.1998:250.
  = Echinocereus knippelianus var. reyesii.
Echinocereus knippelianus var. robustispinus* Hort. nom. nud.
  = Echinocereus knippelianus var. reyesii.
Echinocereus knippelianus var. schaetzleii* Nitzschke 1990:43 nom. prov.
Echinocereus knippelianus var. siberi giberia* Heyer 1989:26 nom. nud.
Echinocereus koehresianus* Hort. ex Taylor 1988:80 pro syn.
  = Echinocereus scheeri var. koehresianus.
Echinocereus koehresii* Hort. ex Taylor 1988:80 pro syn.
  = Echinocereus scheeri var. koehresianus.
Echinocereus koehresii (Frank) Rischer 2009:73
  = Echinocereus scheeri var. koehresianus.
Echinocereus koehresianus subsp. gruberianus W. Rischer 2014. - EcJ-Online 2 (1): 9.
  Type - locality: Mexico, Durango, Municipio Tayoltita; Chalet 5245, 7 May 2010 (IBUG).
Echinocereus krausei De Smet ex Rumpler 1886:789.
  Type - locality: unknown; ? (?).
    Note: This plant is usually listed as a taxonomic synonym of Echinocereus coccineus.
Echinocereus kroenleinii* Hort. nom. nud. = Wilcoxia kroenleinii.
Wilcoxia kroenleinii Cartier 1980:2-3.
  Type - locality: unknown, Kuenzler 379 (P) [Probably from Coahuila, Mexico] .
    Nomenclatural synonyms:
      Echinocereus poselgeri subsp. kroenleinii (Cartier) Lange 1995:138.
      Echinocereus poselgeri fa. kroenleinii* Parker 1996:6 nom. nud.
Echinocereus kuenzleri Castetter et al. 1976:77-78.
  Type - locality: New Mexico, Otero Co., Sacramento Mtns., Elk; Kuenzler 3585, 5 May 1968 (UNM 55571).
    Nomenclatural synonyms:
      Echinocereus pseudohempelii* Kunzler Cat. nom. nud.
      Echinocereus fendleri var. kuenzleri (Castetter et al.) Benson 1982:942.
      Echinocereus fendleri var. hempelii fa. kuenzleri* Parker 1996:2 nom. nud.
      Echinocereus fendleri subsp. kuenzleri* Blum 1997 Seed List nom. nud.
      Echinocereus fendleri subsp. kuenzleri (Castetter et al.) W. Blum & D. Felix. 2014 - EcJ-Online 2 (2): 105.
Echinocereus kuenzlerioides* Hort. nom. nud.
  = Echinocereus kuenzleri.
Echinocereus kunzei Gurke 1907:103-104.
  Type - locality: Arizona, near Phoenix [southern New Mexico according to Britton & Rose 1922:31] ; Kunze (?).
    Nomenclatural synonyms:
      Echinocereus coccineus var. kunzei (Gurke) Backeberg 1960:2070.
        Note: This plant may be a natural hybrid. (see Zimmermann 1993).
Echinocereus laboretianus Daul 1890:76.
  Type - locality: unknown; ? (?).
    Note:Usually considered a synonym of Echinocereus viridiflorus.
Echinocereus labouretianus* Lemaire 1868:57 nom. nud.
  [Probably = Echinocereus laboretianus].
Echinocereus labouretianus var. chrysacanthus* Haage 1900:159 nom. nud.
Echinocereus labouretianus var. gracilispinus* Haage 1900:159 nom. nud.
Echinocereus labouretianus var. sanguineus* Haage 1900:159 nom. nud.
Echinocereus labouretii* Forster ex Haage 1859:19 nom. nud.
Echinocereus labouretii* Forster ex Rumpler 1886:811 pro syn.
  = Echinocereus viridiflorus.
Ecinocereus laboureti var. gracilispinus* Rother 1925:114.
*Echinocereus lamprochlorus = Echinopsis lamprochlora.
Echinocereus laui Frank 1978:74-77.
  Type - locality: Mexico, Sonora, east of Yecora, El Trigo Ranch, 1800m; A. Lau 780, Jan. 1974 (ZSS AA18- 24).
Echinocereus ledingii Peebles 1936:35.
  Type - locality: Arizona, Graham Co., Mt. Graham, 4500 ft.; Louis Wankum, 11 July 1935 (US 1634004).
    Nomenclatural synonyms:
      Echinocereus fendleri var. ledingii (Peebles) Taylor 1985a:253.
Echinocereus leeanus* Lemaire 1868:57 nom. nud.
  [Probably = Echinocereus leeanus Lemaire ex Rumpler].
Echinocereus leeanus (Hooker) Lemaire ex Rumpler 1886:828.
  basionym: Cereus leeanus Hooker 1849:t4417.
    Type - locality: Northern Mexico; ? (?).
      Nomenclatural synonyms:
        Echinocereus polyacanthus var. leeanus* Parker 1996:4 nom. nud.
          Note: Often considered a taxonomic synonym of Echinocereus polyacanthus.
Echinocereus leeanus var. multicostatus (Cels ex Rumpler) Schumann 1897:289.
  = Echinocereus multicostatus.
Echinocereus leonensis Mathsson 1891:66.
  Type - locality: Mexico, Nuevo Leon, near Monterrey; 1800ft. Runge (?).
    Nomenclatural synonyms:
      Echinocereus flaviflorus* Hildmann ex Schumann 1897:264 pro syn.
      Cereus leonensis (Mathsson) Berger 1905:80.
      Echinocereus blanckii var. leonensis (Mathsson) Backeberg1960:1999.
      Echinocereus pentalophus var. leonensis (Mathsson) Taylor 1985b:78.
      Echinocereus pentalophus subsp. leonensis (Mathsson) Taylor 1997:9.
Cereus leptacanthus* De Candolle ex Pfeiffer 1837:101 pro syn.
  = Echinocereus pentalophus var. subarticulatus.
Echinocereus leptacanthus Gaillard 1864:209.
  = Cereus leptacanthus De Candolle ex Salm Dyck 1845:27. nomina nova for Cereus pentalophus var. subarticulatus De Candolle.
Cereus leptacanthus var. crassior* De Candolle ex Forster 1846:373 nom. nud.
Echinocereus leptophus* Gassner Hauptverzeichnis 1937:6 nom. nud.
Echinocereus leucanthus Taylor 1985b:136-137 [New name needed because Echinocereus albiflorus already used].
  basionym: Wilcoxia albiflora Backeberg 1952:16.
    Type - locality: unknown [Probably northwest Sinaloa, Mexico]. (P) [cultivated material from Les Cedres].
Echinocereus leucodictius* Engelmann ex Haage 1859:19 nom. nud.
Echinocereus liebnerianus* Liebner ex Schumann 1896b:262 nom. nud. [Probably a printing error].
  = Echinocereus knippelianus.
*Echinocereus limensis = Haageocereus sp .
Echinocereus lindsayi Meyran 1975:80-83.
  Type - locality: Mexico, Baja Calif. Norte, between Catavina & Laguna Chapala; Meyran et al., 8 May 1975 (MEXU).
    Nomenclatural synonyms:
      Echinocereus ferreirianus var. lindsayi (Meyran) Taylor 1985b:46.
      Echinocereus ferreirianus subsp. lindsayi (Meyran) Taylor 1997:9
Echinocereus llanuraensis (Rutow) W. Blum & D. Felix 2002:3
  = Echinocereus nicholii (Benson) subsp. llanuraensis Rutow
Echinocereus lloydii Britton & Rose 1922:37-38.
  Type - locality: Texas, Pecos Co., Tuna Springs; F.E. Lloyd, 26-28 Feb. 1909 (US 691964).
    Nomenclatural synonyms:
      Echinocereus roetteri var. lloydii (Britton & Rose) Backeberg 1960:2027.
      Echinocereus coccineus fa. lloydii* Parker 1996:4 nom. nud.
      ×Echinocereus lloydii.
        Note: This species is a natural hybrid.
Echinocereus longisetus* Lemaire 1868:57 nom. nud.
  [Probably = Echinocereus longisetus Engelmann ex Rumpler].
Echinocereus longisetus (Engelmann) Engelmann ex Rumpler 1886: 822.
  basionym: Cereus longisetus Engelmann 1856:280. [see Engelmann 1859:32-33, t. 45].
    Type - locality: Mexico, Coahuila, Mtns. near Santa Rosa [Melchor Muzquiz]; J. M. Bigelow, 1853 (MO).
Echinocereus longisetus var. albatus* Sterk 1980:12-13 nom. inval.
  = Echinocereus albatus* Backeberg nom. inval.
Echinocereus longisetus var. delaetii (Gurke) Taylor 1988:79.
  = Echinocereus delaetii.
Echinocereus longisetus subsp. delaetii (Gurke) Taylor 1997:9.
  = Echinocereus delaetii.
Echinocereus longisetus subsp. freudenbergeri (Frank) Blum in Blum et al. 1998a:6. [see Blum et al. 1998b:187].
  = Echinocereus freudenbergeri.
Echinocereus longispinus* Hort. ex Schelle 1926:180 pro syn.
  = Echinocereus acifer.
Echinocereus longispinus Lahman 1950:128. [Originally published Lahman 1936:15 without Latin diagnosis].
  Type - locality: Oklahoma, Mt. Scott; Lahman (MO).
    Note: Usually considered a taxonomic synonym of Echinocereus baileyi.
Echinocereus longispinus var. albispinus* Backeberg 1960:2010 pro syn.
  = Echinocereus albispinus Lahman.
Echinocereus lowryi* Lowry 1936:20 nom. nud.
  [Probably a form of Echinocereus enneacanthus].
Echinocereus luteus Britton & Rose 1913:239.
  Type - locality: Mexico, Sonora, Mtns. above Alamos; Rose et al. 15207, 19 March 1910 (US 535975).
    Nomenclatural synonyms:
      Echinocereus subinermis var. luteus (Britton & Rose) Knuth in Backeberg & Knuth 1935:312.
      Echinocereus subinermis fa. luteus* Parker 1996:4 nom. nud.
Echinocereus madrensis Patoni 1910:42.
  Type - locality: Mexico, Durango, high Sierra Madre; ? (?).
    Nomenclatural synonyms:
      Echinocereus adustus var. schwarzii fa. madrensis* Parker 1996:6. nom. nud.
Echinocereus malibrani* Rebut ex Haage 1892:118 nom. nud.
  [Probably = Echinocereus dubius (According to Schelle 1907:135) ].
Echinocereus mamillatus (Engelmann ex Coulter) Britton & Rose 1922:41.
  basionym: Cereus mamillatus Engelmann ex Coulter 1896:405.
    Type - locality: Mexico, Baja California, south of Mulege, Gabb 16, 1867 (MO).
      Nomenclatural synonyms:
        Echinocereus brandegeei fa. mamillatus* Parker 1996:1 nom. nud.
          Note: Usually considered a taxonomic synonym of Echinocereus brandegeei.
Echinocereus mamillifer* Forster ex Haage 1859:19 nom. nud.
Echinocereus mamilliter* Forster ex Seitz 1870:11 nom. nud.
Echinocereus mamillosus Hort. ex Rumpler 1886:787.
  Type - locality: unknown.
Echinocereus mapimiensis Anderson et al. 1998:284-285.
  Type - locality: Mexico, Coahuila, Bolson de Mapimi, Keil 8136, 22 Aug 1971 (DES).
Echinocereus marginii* Ely 1935:6 nom. nud.
Echinocereus mariae* Backeberg 1965:2, 19 nom. inval. (no type designated).
  Note: A form of Echinocereus baileyi from northwest of Granite, Oklahoma.
Echinocereus maritimus (Jones) Schumann 1897:273-274.
  basionym: Cereus maritimus Jones 1883:973.
    Type - locality: Mexico, Baja California, Norte, near Ensenada; M.E. Jones, April 1882 (POM).
      Nomenclatural synonyms:
        Cereus glomeratus* Orcutt 1897:21 pro syn.
          Note: Echinocereus orcuttii and Echinocereus flaviflorus Coulter ex Schumann are usually considered taxonomic
            synonyms of this species.
Echinocereus maritimus var. hancockii Taylor 1985b:44
  = Echinocereus hancockii.
Echinocereus maritimus subsp. hancockii (Dawson) Blum & Rutow in Blum et al. 1998a:6. [see Blum et al. 1998b:57].
  = Echinocereus hancockii.
Echinocereus marksianus* Schwarz ex Backeberg 1966:124 nom. inval. (no type designated).
  Note: From Northern Mexico. Taylor (1985b) lists this name as a synonym of Echinocereus polyacanthus var. densus.
Echinocereus matlesianus* Backeberg ex Bravo 1991:26 misspelling = Echinocereus matthesianus* Backeberg.
Echinocereus matthesianus* Backeberg 1963:6 nom. inval. (no type designated).
  Note: Taylor (1993) lists this name as a synonym of Echinocereus polyacanthus.
Echinocereus matudae Bravo 1961:119-121.
  Type - locality: Mexico, Chihuhaua, Rio Papigochic near Temosochic; Matuda (MEXU).
    Nomenclatural synonyms:
      Echinocereus arizonicus subsp. matudae (Bravo) Rutow in Lange & Rutow 1994:51.
      Echinocereus coccineus var. arizonicus fa. matudae* Parker 1996:4 nom. nud.
Echinocereus melaleuca* Foster ex Haage 1892:117 nom. nud.
  [Probably= Echinocereus fendleri].
Echinocereus melanacanthus* Engelmann ex Earl 1963:61 printing error.= Echinocereus coccineus.
Echinocereus melanocentrus* Lowry 1936:20 nom. inval. (no Latin diagnosis).
  = Echinocereus melanocentrus* Lowry ex Backeberg.
Echinocereus melanocentrus* Lowry ex Backeberg 1960:2030 nom. inval. (no type designated).
  Note: usually considered a synonym of Echinocereus reichenbachii var. albertii.
Echinocereus merkeri* Hildmann ex Haage 1892:118 nom. nud.
Echinocereus merkeri Hildmann ex Schumann 1897:277.
  Type - locality: unknown [range given as Durango near Cd. Lerdo, Coahuila, Sierra Bola & near Parras].
    Nomenclatural synonyms:
      Echinocereus jacoybi* Hort. ex Schumann 1897:278 pro syn.
      Cereus merkeri (Hildmann ex Schumann) Berger 1905:81.
      Echinocereus enneacanthus fa. merkeri* Parker 1996:2 nom. nud.
        Note: Taylor (1985b) lists this species as a taxonomic synonym of Echinocereus enneacanthus.
Echinocereus merkeri var. densispinus* Schmoll ex Backeberg 1960:2051 nom. nud.
Echinocereus metornii Frank 1990b:210-218.
  Type - locality: Mexico, Coahuila, Sierra Mojada, 1500m, Metorn 49, 4 April 1989 (ZSS).
    Nomenclatural synonyms:
      Echinocereus chisoensis var. metornii* Parker 1996:5 nom. nud.
      Echinocereus fobeanus subsp. metornii (Frank) Blum & Lange in Blum et al. 1998b:307.
        Note: Taylor (1993) lists this species as a taxonomic synonym of Echinocereus fobeanus.
Echinocereus milleri Blum, Kunzler & Oldach. in Blum & Oldach 1999:69-76.
  Type - locality: Robert Lee, Coke Co., Texas. Leuck 229, August 1978 (OKL).
    Nomenclatural synonyms:
      Echinocereus russanthus subsp. milleri* Blum 1997 Seed List nom. nud.
Echinocereus moelleri* Hort. (Cactus & Succulents Journal (US) 13:22 1941) nom. nud.
Echinocereus mojavensis (Engelmann & Bigelow) Engelmann ex Rumpler 1886:803.
  basionym: Cereus mojavensis Engelmann & Bigelow in Engelmann 1856:281.[see Engelmann 1857:33].
    Type - locality: California, Mojave Creek; J.M. Bigelow, 4 March 1854 (MO).
      Nomenclatural synonyms:
        Cereus bigelovii* Engelmann 1856:pl. 4 nom. nud.
        Echinocereus triglochidiatus var. mojavensis (Engelmann & Bigelow) Benson 1944:255.
        Echinocereus coccineus var. mojavensis* Parker 1996:4 nom. nud.
        Echinocereus coccineus subsp. mojavensis (Engelmann & Bigelow) Taylor 1997:8.
        Echinocereus triglochidiatus subsp. mojavensis (Engelmann & Bigelow) Blum & Lange in Blum et al. 1998b:362.
Echinocereus mojavensis var. albispinus* Hort. ex Schelle1926:167 nom. nud.
Echinocereus mojavensis fa. inermis* Mesa Garden Seed Catalog1996 nom. nud.
  = Echinocereus coccineus var. inermis.
Echinocereus mojavensis var. zuniensis (Engelmann & Bigelow) Engelmann ex Rumpler 1886:803.
  basionym: Cereus mojavensis var. zuniensis Engelmann & Bigelow in Engelmann 1856:281.
    Type - locality: Arizona, Canon Diablo, on Little Colorado River; J.M. Bigelow, 14 Dec. 1853 (MO).
      Nomenclatural synonyms:
        Cereus bigelovii var. zuniensis* Engelmann 1857: t. 4 nom. nud.
        Echinocereus zuniensis* Eng. & Big. ex Haage & Schmidt nom. nud.
Echinocereus mombergerianus Frank 1990c:261 (Originally published Frank 1989:272-277 with no type designated).
  Type - locality: Mexico, Baja California Norte, Sierra San Pedro Martir, Hidalgo Observatory; Lau 1248 (ZSS).
    Nomenclatural synonyms:
      Echinocereus polyacanthus var. mombergerianus* Parker1996:4 nom. nud.
      Echinocereus polyacanthus subsp. mombergerianus (Frank) Breckwoldt & Matylewicz 1996:15.
      Echinocereus pacificus subsp. mombergerianus Blum, Rischer & Rutow in Blum et al. 1998b:418.
Echinocereus monacanthus Heese 1904:215.
  Type - locality: Texas - Mexico Border, Rancho Tacubaya.
    Note: Usually considered a taxonomic synonym of Echinocereus paucispinus.

Echinocereus morricalii – Echinocereus reichenbachianus

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Echinocereus morricalii Riha 1975:75.
  Type - locality: Mexico, Nuevo Leon, Barranca de las Garrapatas south of Monterrey; D. B. Morrical (PRC 322).
    Nomenclatural synonyms:
      Echinocereus viereckii var. morricalii (Riha) Taylor 1985b:93.
      Echinocereus blanckii var. inermis* Heyer 1989:26 pro syn.
      Echinocereus viereckii subsp. morricalii (Riha) Taylor 1997:10.
*Echinocereus multiangularis = Haageocereus sp.
*Echinocereus multiangularis var. limensis = Haageocereus sp.
*Echinocereus multiangularis var. maletianus = ?
*Echinocereus multiangularis var. pallidior = ?
Echinocereus multicostatus Cels ex Rumpler 1886:834.
  Type - locality: unknown; ? (?).
    Nomenclatural synonyms:
      Echinocereus leeanus var. multicostatus (Cels ex Rumpler) Schumann 1897:289.
      Echinocereus triglochidiatus var. multicostatus (Cels ex Rumpler) Marshall 1953:67.
        Note: this species is often considered a taxonomic synonym of Echinocereus leeanus, but its may be of South American
          origin and not an Echinocereus.
Echinocereus munzii (Parish) Benson 1941:361.
  basionym: Cereus munzii Parish 1926:48.
    Type - locality: California, Riverside Co., 2 miles below Kenworthy, 1400m.; Munz & Johnson 5570, 21 May 1922 (UC).
      Nomenclatural synonyms:
        Echinocereus engelmannii var. munzii (Parish) Pierce & Fosberg 1933:123.
        Echinocereus engelmannii fa. munzii* (Parish) Ruinaard, Succulenta 89(3): , Nom. inval. (Art. 33.4),
Echinocereus neocapillus (Weniger) Blum & Lange in Blum et al. 1998b:219.
  = Echinocereus chloranthus var. neocapillus.
Echinocereus neomexicanus Standley 1908:87-88.
  Type - locality: New Mexico, Dona Ana Co., mesa west of Organ Mtns.; Standley 383, 1907 (MO).
    Nomenclatural synonyms:
      Echinocereus triglochidiatus var. neomexicanus (Standley) Standley ex Marshall & Bock 1941:118.
      Cereus neomexicanus (Standley) Tidestrom 1941:298.
      Echinocereus polyacanthus var. neomexicanus* Weniger1970:44 nom. inval.
      Echinocereus coccineus var. neomexicanus* Parker 1996:4 nom. nud.
      × Echinocereus neomexicanus Blum et al. 1998b:449.
      ×Echinocereus roetteri var. neomexicanus ( J.M.Coult. ) A. D. Zimmerman in Powell & Weedin 2004:211. pro sp. hybr.
        Note: This plant has been confused with Echinocereus rosei under the name Echinocereus triglochidiatus var. neomexicanus
          [Benson] and Echinocereus polyacanthus [Bravo 1991:24]. It is probably a natural hybrid of E. rosei and E. chloranthus.
Echinocereus neoroemeri* (Muhlenpfordt) Ito 1971:227. nom. illegit.
  = Echinocereus roemeri (Muhlenpfordt) Rydberg.
Wilcoxia nerispina* Backeberg 1960:2078 nom. invalid.
  = Echinocereus schmollii.
Echinocereus nicholii (Benson) Parfitt 1987:157-158.
  = Echinocereus engelmannii var. nicholii.
Echinocereus nicholii fa. guaymasensis* Parker 1996:1 nom. nud.
Echinocereus nicholii subsp. llanuraensis Rutow 1995:61-71.
  Type - locality: Mexico, Sonora, Guaymas 300m; (ZSS  A 16936).
    Nomenclatural synonyms:
      Echinocereus nicholii fa. llanuraensis* Parker 1996:1 nom. nud.
      Echinocereus engelmannii subsp. llanuraensis ( Rutow ) Felger 1999: 20.
Echinocereus nicholii fa. llanuraensis* Parker 1996:1 nom. nud.
  = Echinocereus nicholii subsp. llanuraensis.
Echinocereus nigrispinus* Kohres Catalog 1979. nom. nud.
Echinocereus nivilus* Heyer 1989:26 nom. nud. [= Lau 1101].
Echinocereus nivosus Glass & Foster 1978a:18-19.
  Type - locality: Mexico, Coahuila, Southwest of General Cepeda, 3 km. north of El Cinco; Glass & Foster 3764, Feb. 1972
    (POM).
      Note: Echinocereus albatus* Backeberg 1960:2007 nom. inval. is usually considered a taxonomic synonym of this species.
Echinocereus nivosus fa. albatus* Parker 1996:3 nom. nud.
  = Echinocereus albatus*.
Echinocereus noctiflorus* Hort. ex Backeberg 1960:1991 pro syn.
  = Echinocereus cucumis.
Echinocereus nocturniflorus* Hort. ex Backeberg 1960:1991 pro syn.
  = Echinocereus cucumis.
Echinocereus nogalensis* Haage 1900:159 nom. nud. [Perhaps = Echinocereus pectinatus var. rigidissimus].
Echinocereus occidentalis (Taylor) Richer et al. 2009:23.
  = Echinocereus stramineus var. occidentalis Taylor 1988:70-71.
Echinocereus ochoterenae Ortega 1928:88-90.
  Type - locality: Mexico, Sinaloa, near Concordia, Cerro de la Cobriza; J.G. Ortega, 24 May 1927 (MEXU?).
    Nomenclatural synonyms:
      Echinocereus subinermis var. ochoterenae (Ortega) Unger 1984:164.
      Echinocereus subinermis subsp. ochoterenae (Ortega) Taylor 1997:10.
        Note: Ortega described varieties "A" and "B", but did not give them names.
Echinocereus octacanthus (Muhlenpfordt) Britton & Rose 1922:13.
  basionym: Echinopsis octacantha Muhlenpfordt 1848:19.
    Type - locality: Northern Texas [San Saba River according to Engelmann]; Roemer (?).
      Nomenclatural synonyms:
        Cereus octacanthus (Muhlenpfordt) Coulter 1896:395.
        Echinocereus coccineus var. octacanthus* Boissevain & Davidson 1940:39 nom. illegit.
        Echinocereus triglochidiatus var. octacanthus* Muhlenpfordt ex Marshall & Bock 1941:118. nom. illegit.
          Note: The proper application of this name is uncertain. If it can be shown to be one of the members of the Echinocereus
            coccineus group it would have priority since it was published first. This would cause numerous nomenclatural changes.
Echinocereus occidentalis (Taylor) Rischer & Breckwoldt 2009:23.
  = Echinocereus stramineus subsp. occidentalis Taylor 1997:10.
Echinocereus oklahomensis Lahman 1950:128. [Originally published in Lahman 1935:141 without Latin diagnosis].
  Type - locality: Oklahoma, Wichita Mtns.; Lahman (MO).
    Note: Usually consisdered a taxonomic synonym of Echinocereus baileyi.
Echinocereus orcuttii Rose ex Orcutt 1926:4.
  Type - locality: Mexico, Baja California, Cape Colset; Orcutt (?).
    Note: Usually consisdered a taxonomic synonym of Echinocereus maritimus.
Echinocereus ortegae Rose ex Ortega 1929:24.
  Type - locality: Mexico, Sinaloa, Cuenca del Rio Tamazula, Sianori, 800m.; Ortega (?).
    Note: Taylor (1993) lists Echinocereus scheeri var. koehresianus as a taxonomic synonym of this species.
Echinocereus ortegae subsp. koehresianus (Frank) Rischer & Frank 1996:93.
  = Echinocereus scheeri var. koehresianus.
Echinocereus pacificus* Haage 1892:118 nom.nud.
Echinocereus pacificus (Engelmann ex Orcutt) Britton & Rose 1922:12.
  basionym: Cereus phoeniceus var. pacificus Engelmann ex Orcutt 1886:46.
    Type - locality: Mexico, Baja California Norte, Rio San Carlos; Parry et al., 25 Jan. 1883 (MO).
      Nomenclatural synonyms:
        Cereus pacificus (Engelmann ex Orcutt) Coulter 1896:397.
        Echinocereus triglochidiatus var. pacificus (Engelmann ex Orcutt) Engelmann ex Marshall & Bock 1941:118.
        Echinocereus polyacanthus var. pacificus (Engelmann ex Orcutt) Taylor 1984:160.
        Echinocereus polyacanthus subsp. pacificus (Engelmann ex Orcutt) Breckwoldt 1996:15.
Echinocereus pacificus subsp. mombergerianus Blum, Rischer & Rutow in Blum et al. 1998b:418.
  = Echinocereus mombergerianus.
Echinocereus palmeri Britton & Rose 1922:34.
  Type - locality: Mexico, Chihuahua, hill near Cd. Chihuahua; E. Palmer 121, Apr. 1908 (US).
Echinocereus palmeri subsp. escobedensis Rischer 2003b:54.
  Type - locality: Mexico, Durango; Lau 1565, 18 March 1988 (PR 11691a) ex cult. coll. Rischer.
Echinocereus palmeri subsp. mazapil H.M.Hern. & Gomez-Hin.2004:2.
  Type - locality: Mexico, Zacatecas, Municipio Mazapil; H.M. Hernández, C. Gómez-Hinostrosa & B. Goettsch 3528 (MEXU).
Echinocereus pamanesiorum Lau 1981:36-41.
  Type - locality: Mexico, Zacatecas, Rio Huaynamota, bridge between Huejuquilla & San Juan Capistrano; Lau 1247 (MEXU).
Echinocereus pamanesiorum subsp. bonatzii (Romer) Romer 1997:45.
  = Echinocereus bonatzii.
Echinocereus papillosus Linke ex Rumpler 1886:783.
  Type - locality: unknown; ? (?).
    Nomenclatural synonyms:
      Cereus papillosus (Linke ex Rumpler) Berger 1905:80.
      Echinocereus blanckii var. papillosus (Linke ex Rumpler) Benson 1969d:126.
      Echinocereus berlandieri var. papillosus (Linke ex Rumpler) Benson 1976:59.
        Note: Benson uses the names Echinocereus blanckii var. angusticeps and Echinocereus berlandieri var. angusticeps for this
          plant. Usually listed as Echinocereus papillosus Linke ex Rumpler 1886:783.
Echinocereus papillosus var. angusticeps (Clover) Marshall & Bock 1941:119.
  = Echinocereus angusticeps.
Echinocereus papillosus subsp. angusticeps* Blum 1997 Seed List nom. nud.
  = Echinocereus angusticeps.
Echinocereus papillosus var. giganteus* Bailey 1941:270 nom. nud.
Echinocereus papillosus var. rubescens Hildmann ex Schelle1926:165.
  Type - locality: unknown; ? (?).
Echinocereus parkeri Taylor 1988:73.
  Type - locality: Mexico, Nuevo Leon, between La Poza & Puerto de los Pastores; Hansen et al. 3863 (MEXU).
    Nomenclatural synonyms:
      Echinocereus stramineus var. parkeri* Seed list 1996 - British Cactus and Succulent Journal. 
Echinocereus parkeri subsp. arteagensis Blum & Lange in Blum et al. 1998b:175.
  Type - locality: Mexico, Coahuila - Nuevo Leon Border, Arteaga Canyon, 2250 m.; U. Guzman in Sánchez- Mejorada 3807
    (MEXU).
    Nomenclatural synonyms:
      Echinocereus parkeri subsp. arteagensis* (Blum & Lange) Frank & Scherer 1998:61. nom. inval.
Echinocereus parkeri subsp. arteagensis* (Blum & Lange) Frank & Scherer 1998:61. nom. inval.
Echinocereus parkeri fa. barcena* Parker 1996:3 nom. nud.
  = Echinocereus barcena.
Echinocereus parkeri var. gonzalezii Taylor 1988:74.
  Type - locality: Mexico, Tamaulipas, Sierra de las Vacas, 4km NE of San Jose d'Llano 1750m.; Gonzalez Medrano 8515 (MEXU).
Echinocereus parkeri subsp. gonzalezii (Taylor) Taylor 1997:9.
  = Echinocereus parkeri var. gonzalezii.
Echinocereus parkeri var. mazapilensis* K. Neumann 1993 nom. prov. Kaktusblüte p. 55.
Echinocereus parkeri subsp. mazapilensis Blum & Lange in Blum et al. 1998b:172.
  Type - locality: Mexico, Zacatecas, 0.3 km from El Cobre near Mazapil, 2750 m.; Sanchez - Mejorada et al. 4041 (MEXU).
Echinocereus parkeri subsp. rayonesensis (Taylor) Felix & Blum 2011:99.
  = Echinocereus rayonesensis Taylor 1988:75-76.
Echinocereus paucispinus (Engelmann) Loder 1882:445.
  basionym: Cereus paucispinus Engelmann 1856:285. [see Engelmann 1859:34].
    Type - locality: Texas, San Pedro River; Wright, 1849 (MO). cultivated from original seeds.
      Nomenclatural synonyms:
        Echinocereus paucispinus* Engelmann ex Rumpler 1886:794 nom. illegit.
        Echinocereus triglochidiatus var. paucispinus (Engelmann) Engelmann ex Marshall & Bock 1941:117.
        Echinocereus coccineus var. paucispinus (Engelmann) Ferguson 1989:222.
        Echinocereus coccineus subsp. paucispinus Blum, Lange & Rutow in Blum et al. 1998b:433.
          Note:Usually cited as Echinocereus paucispinus Engelmann ex Rumpler 1886:794.
Echinocereus paucispinus var. flavispinus* Haage 1892:118 nom. nud.
Echinocereus paucispinus var. flavispinus* Walton 1899b:18 nom. nud.
Echinocereus paucispinus var. flavispinus Hort. ex Schelle 1926:178.
  Type - locality: unknown; ? (?).
Echinocereus paucispinus var. fulvispinus* Haage 1897:12 nom. nud.
Echinocereus paucispinus var. fulvispinus Bowles 1908:28.
  Type - locality: unknown; ? (?).
Echinocereus paucispinus var. gonacanthus (Engelmann & Bigelow) Schumann 1897:281
  = Echinocereus gonacanthus.
Echinocereus paucispinus var. hexaedrus (Engelmann & Bigelow) Schumann 1897:281
  = Echinocereus hexaedrus.
Echinocereus paucispinus var. nigrispinus* Haage 1897:12 nom. nud.
Echinocereus paucispinus var. nigrispinus Bowles 1908:28.
  Type - locality: unknown; ? (?).
Echinocereus paucispinus var. triglochidiatus* Schumann 1897:281 nom. illegit.
  = Echinocereus triglochidiatus.
Echinocereus pectinatus (Scheidweiler) Engelmann 1848a:110.
  basionym: Echinocactus pectinatus Scheidweiler 1838:492.
    Type - locality: Mexico, San Luis Potosi, Villa del Penasco; Galeotti (?).
      Nomenclatural synonyms:
        Echinopsis pectinata (Scheidweiler) Fennel 1843:282.
        Cereus pectinatus (Scheidweiler) Engelmann 1849:50.
          Note: Echinocereus cernosus* Frič 1929:24 nom. nud. is listed by Backeberg as a synonym of this species.
            Echinocactus pectiniferus Lemaire 1839:25 is usually considered a taxonomic synonym of this species.
Echinocereus pectinatus var. adustus* Haage 1859:19 nom. nud.
Echinocereus pectinatus var. adustus [mispelled adjustus] (Engelmann) Daul 1890:78.
  = Echinocereus adustus.
  Note: Usually ascribed to Schumann 1897:271.
Echinocereus pectinatus fa. adustus (Engelmann) Voss 1894:378.
  = Echinocereus adustus.
Echinocereus pectinatus fa. adustus subfa. castaneus (Engelmann) Engelm. ex Schelle 1907:132.
Echinocereus pectinatus var. adustus fa. castaneus (Engelmann) Engelm. ex Schelle 1926:174.
  = Echinocereus caespitosus var. castaneus.
Echinocereus pectinatus var. armatus* Engelmann ex Seitz 1870:11 nom. nud.
Echinocereus pectinatus var. armatus (Poselger) Poselger ex Rumpler 1886:818.
  basionym: Cereus pectinatus var. armatus Poselger 1853:134.
    Type - locality: Mexico, Nuevo Leon, near Monterrey; Poselger (?).
      Nomenclatural synonyms:
        Cereus pectinatus var. spinosus* Coulter 1896:357.
        Echinocereus spinosus Coulter ex Schelle 1926:174 pro syn.
        Echinocereus armatus (Poselger) Poselger ex Backeberg & Knuth 1935:318.
        Echinocereus caespitosus var. armatus (Poselger) Poselger ex Borg 1937:174.
        Echinocereus reichenbachii var. armatus (Poselger) Taylor 1985b:133.
        Echinocereus fitchii var. armatus* Mesa Garden Seed Catalog 1996 nom. nud.
        Echinocereus reichenbachii subsp. armatus (Poselger) Taylor 1997:9.
        Echinocereus fitchii subsp. armatus (Poselger) Blum et al. 1998b:314.
Echinocereus pectinatus var. bauerii Rother 1923:127.
  = Echinocereus bauerii Rother 1902:61
Echinocereus pectinatus var. bristolii (Marshall) Marshall 1956b:81-82.
  = Echinocereus bristolii.
Echinocereus pectinatus var. caespitosus (Engelmann) Schumann 1897:272.
  = Echinocereus caespitosus.
Echinocereus pectinatus fa. candicans* Hort. ex Schelle 1907:132 nom. nud.
Echinocereus pectinatus var. candidissimus* Seitz 1878:11 nom. nud.
Echinocereus pectinatus var. castaneus* Seitz 1878:11 nom. nud.
Echinocereus pectinatus var. castaneus (Engelmann) Daul 1890:78.
  = Echinocereus caespitosus var. castaneus.
Echinocereus pectinatus fa. castaneus (Engelmann) Krainz 1967:50.
  = Echinocereus caespitosus var. castaneus.
*Echinocereus pectinatus var. centralis Coulter ex Schumann 1897:271.
  error in determination. = Echinomastus centrilis.
Echinocereus pectinatus var. chloranthus* Haage 1892:118 nom. nud.
Echinocereus pectinatus var. chrysacanthus Schumann 1897:272.
  Type - locality: Mexico; ? (?).
    Nomenclatural synonyms:
      Echinocereus caespitosus var. chrysacanthus (Schumann) Schumann ex Borg 1937:174.
Echinocereus pectinatus var. crassispinus Runge ex Mathsson 1890:465.
  Type - locality: Mexico, Coahuila, Saltillo; Mathsson (?).
Echinocereus pectinatus var. ctenoides* Hort. ex Schelle1907:130 pro syn.
  = Echinocereus ctenoides.
Echinocereus pectinatus var. ctenoides* Weniger 1971:31 nom. inval. (no basionym given).
  = Echinocereus ctenoides.
Echinocereus pectinatus var. ctenoides (Engelmann) Weniger ex Frank 1991:135.
  = Echinocereus ctenoides.
Echinocereus pectinatus subsp. ctenoides (Engelmann) Frank 1997:136.
  = Echinocereus ctenoides.
Echinocereus pectinatus var. dasyacanthus (Engelmann) Engelmann ex Haage 1892:118.
  = Echinocereus dasyacanthus.
Echinocereus pectinatus var. dasyacanthus* Earle 1971:80 nom. illegit. (later homonym).
  = Echinocereus dasyacanthus.
Echinocereus pectinatus var. dasyacanthus* Taylor 1984:179 nom. illegit. (later homonym).
  = Echinocereus dasyacanthus.
Echinocereus pectinatus var. grandicostatus* Schmoll ex Backeberg 1960:2043 nom. nud.
Echinocereus pectinatus var. grandis (Britton & Rose) Marshall 1955:68.
  = Echinocereus grandis.
Echinocereus pectinatus var. jamauvei* Frič ex Orcutt 1926:4 pro syn.
  = Echinocereus castaneus.
Echinocereus pectinatus var. labouretianus* Seitz 1870:11 nom. nud.
Echinocereus pectinatus var. laevior* (Monatsschrift fur Kakteenkunde index 1912:56) nom. nud.
Echinocereus pectinatus var. minor (Engelmann) Benson 1968:125 for Echinocereus dasyacanthus var. minor*.
  Note: Benson uses this name for Echinocereus roetteri.
Echinocereus pectinatus var. neomexicanus (Coulter) Benson 1944:256.
  = Echinocereus dasyacanthus var. neomexicanus*.
    Note: Benson uses this name for Echinocereus dasyacanthus.
Echinocereus pectinatus var. ordustus* Lakerveld Catalog. nom. nud.
Echinocereus pectinatus var. pailanus* Schwarz ex Backeberg 1960:2043 nom. nud.
Echinocereus pectinatus var. pailensis* Backeberg 1937:21. nom. nud.
Echinocereus pectinatus var. pailensis* Bailey 1941:271 nom. nud.
Echinocereus pectinatus var. pictus* Seitz 1870:11 nom. nud.
Echinocereus pectinatus var. rectispinus* Zimmerman1993:277 nom. nud.
  = Echinocereus dasyacanthus var. rectispinus.
Echinocereus pectinatus var. reichenbachii (Terscheck ex Walpers) Werdermann 1930:t5.
  = Echinocereus reichenbachii.
Echinocereus pectinatus var. rigidissimus (Engelmann) Engelmann ex Rumpler 1886:818.
  basionym: Cereus pectinatus var. rigidissimus Engelmann 1856:279.
    Type - locality: Arizona, Sierra Pajarito; Schott, 1855 (MO 2016861).
      Nomenclatural synonyms:
        Echinocereus pectinatus var. robustior* Hort. ex Hirscht 1897:95 pro syn.
        Echinocereus candicans* Runge ex Haage 1900:157 pro syn.
        Echinocereus rigidissimus (Engelmann) Hort. ex F. A. Haage 1897:13.
        Echinocereus robustior* Britton & Rose 1922:28 pro syn.
        Echinocereus dasyacanthus var. rigidissimus (Engelmann) Marshall & Bock 1941:119.
        Echinocereus pectinatus fa. rigidissimus (Engelmann) Krainz 1967:50.
          Note: Usually listed as Echinocereus rigidissimus Rose 1909a:293. Echinocereus nogalensis* Haage 1900:159 nom. nud. is
            usually considered a synonym of this plant.
Echinocereus pectinatus fa. rigidissimus (Engelmann) Krainz 1967:50.
  = Echinocereus pectinatus var. rigidissimus.
Echinocereus pectinatus var. robustior* Hort. ex Hirscht 1897:95 pro syn.
  = Echinocereus pectinatus var. rigidissimus.
Echinocereus pectinatus var. robustus Bauer ex Wittmack 1890:513.
  Type - locality: Mexico, Sonora, Nogales; ? (?).
    Note: usually considered a taxonomic synonym of Echinocereus pectinatus var. rigidissimus.
Echinocereus pectinatus var. rotatus (Linke) Linke ex Haage 1892:118.
  = Echinocereus rotatus.
Echinocereus pectinatus var. rubescens* Weniger 1970:21-22 nom. nud.
Echinocereus pectinatus var. rubispinus Frank & Lau in Frank 1982:32-35.
  Type - locality: Mexico, Chihuahua, Canon de Barbarocas in Sierra Obscura, 1600-1800m.; Lau 088 (ZSS AA 18-30).
    Nomenclatural synonyms:
      Echinocereus rigidissimus var. rubispinus (Frank & Lau) Taylor 1984:175.
      Echinocereus rigidissimus subsp. rubispinus Taylor 1997:9.
Echinocereus pectinatus var. rungei* Hort. ex Schelle 1926:174 nom. nud.
Echinocereus pectinatus var. rufispinus* Daul 1890:78 nom. nud.
Echinocereus pectinatus var. rufispinus Schumann 1897:272.
  Type - locality: Mexico; ? (?).
    Nomenclatural synonyms:
      Echinocereus caespitosus var. rufispinus (Schumann) Schumann ex Borg 1951:221.
Cereus pectinatus fa. rufispinus (Engelmann) Voss 1894:378.
  = Echinocereus rufispinus.
E. pectinatus subsp. rutowiorum Blum 2014:24-32.
  Type - locality: Mexico; Chihuahua, Municipio de Chihuahua, east  of El Olivio; Anonymous 356, 2006, 1650m, April 2006
    (KR21723).
Echinocereus pectinatus var. scopulorum (Britton & Rose) Marshall 1955:61.
  = Echinocereus scopulorum.
Echinocereus pectinatus var. sonorensis* (Kaktusy 1973 according to Evanich 1989:111).
Cereus pectinatus var. spinosus* Coulter 1896:357 nom. illegit.
  Note: an unnecessary new name for Echinocereus pectinatus var. armatus.
Echinocereus pectinatus var. steereae* Rumpler ex Weniger 1970:32 pro syn.
  = Echinocereus steereae.
Echinocereus pectinatus var. tamaulipensis Frič ex Schelle 1926:174.
  Type - locality: unknown; ? (?).
    Nomenclatural synonyms:
      Echinocereus caespitosus var. tamaulipensis (Frič) Frič ex Borg 1937:174.
Echinocereus pectinatus var. texensis* Horner ex Seitz 1870:11 nom. nud.
Echinocereus pectinatus var. texensis Hort. ex Rumpler 1886:817.
  Type - locality: Texas; ? (?).
Echinocereus pectinatus fa. texensis* Hooker ex Schelle 1907:132 nom. nud.
Echinocereus pectinatus var. wenigeri Benson 1968:124.
  Type - locality: Texas, Valverde Co, near Langtry, Benson 16,521 (POM 311338).
    Nomenclatural synonyms:
      Echinocereus pectinatus subsp. wenigeri.
Echinocereus pectinatus subsp. wenigeri (Benson) Blum & Rutow in Blum et al. 1998: 111.
  = Echinocereus pectinatus var. wenigeri.
Echinocactus pectiniferus Lemaire 1839:25.
  Type - locality: unknown; ? (?).
    Nomenclatural synonyms:
      Cereus pectiniferus (Lemaire) Labouret 1853:320.
        Note: usually considered a taxonomic synonym of Echinocereus pectinatus.
Echinocactus pectiniferus var. laevior Monville ex Lemaire 1839:26.
  Type - locality: unknown; ? (?).
    Nomenclatural synonyms:
      Echinopsis pectinata var. laevior (Monville ex Lemaire) Monville ex Forster 1846:365.
*Echinocereus pencilliformis Linke 1858 [of Bolivian origin and thus not an Echinocereus].
Echinocereus pensilis (K. Brandegge) J. A. Purpus 1908:5.
  basionym: Cereus pensilis K. Brandegee 1904:192.
    Type - locality: Mexico, Baja California Sur, Sierra de Laguna; T. S. Brandegee 246, Jan/Feb. 1890 (UC).
      Nomenclatural synonyms:
        Morangaya pensilis (K. Brandegee) Rowley 1974b:44-45.
Echinocereus pentalophus* Engelmann ex Haage 1859:20 nom. nud.
Echinocereus pentalophus* Lemaire 1868:56. nom. nud.
  [Probably = Echinocereus pentalophus De Candolle ex Rumpler].
Echinocereus pentalophus (De Candolle) Lemaire ex Rumpler 1886:774.
  basionym: Cereus pentalophus De Candolle 1828:117.
    Type - locality: Mexico, Hidalgo, near Zimapan; Coulter (?).
      Nomenclatural synonyms:
        Cereus propinquus* Hort. Genev. ex Walpers 1843:275 pro syn.
        Echinocereus pentalophus var. aureovariegatus* Ito 1952:134 nom. nud.
Echinocereus pentalophus var. ehrenbergii (Pfeiffer) Backeberg 1960:2003.
  = Echinocereus ehrenbergii.
Echinocereus pentalophus var. leonensis (Mathsson) Taylor 1985b:78.
  = Echinocereus leonensis.
Echinocereus pentalophus subsp. leonensis (Mathsson) Taylor 1997:9.
  = Echinocereus leonensis.
Echinocereus pentalophus var. leptacanthus* Haage 1859:20 nom. nud.
Echinocereus pentalophus var. leptacanthus* Lemaire ex Rumpler 1886:775 nom. illegit.
  = Echinocereus leptacanthus.
Echinocereus pentalophus var. procumbens (Engelmann) Fournier 1935:25.
  = Echinocereus procumbens.
Echinocereus pentalophus fa. procumbens* Parker 1996:4 nom. nud.
  = Echinocereus procumbens.
Echinocereus pentalophus subsp. procumbens (Engelmann) Blum & Lange in Blum et al. 1998b:129.
  = Echinocereus procumbens.
Cereus pentalophus var. radicans DeCandolle1828:117.
  Type - locality: Mexico, Hidalgo, near Zimapan; Coulter (?).
    Nomenclatural synonyms:
      Cereus propinquus* Hort. Genev. ex Walpers 1843:275 pro syn.
Echinocereus pentalophus var. radicans* Weniger 1970:51 nom. inval.
Echinocereus pentalophus var. radicans* Weniger 1970:51 nom. inval. (no basionym given).
  = Cereus pentalophus var. radicans.
Cereus pentalophus var. simplex De Candolle 1828:117.
  Type - locality: Mexico, Hidalgo, near Zimapan; Coulter (?).
    Nomenclatural synonyms:
      Cereus propinquus* Salm Dyck ex Otto:1833:366. nom. nud.
      Cereus propinquus* De Candolle ex Walpers 1843:333 pro syn.
      Echinocereus propinquus De Candolle ex Haage 1892:118.
      Echinocereus pentalophus var. simplex* Weniger 1970:51nom. inval.
Echinocereus pentalophus var. simplex* Weniger 1970:51 nom. inval. (no basionym given).
  = Cereus pentalophus var. simplex.
Cereus pentalophus var. subarticulatus De Candolle 1828:117.
  Type - locality: Hidalgo, near Zimapan; Coulter (?).
    Nomenclatural synonyms:
      Cereus propinquus var. subarticulatus* Pfeiffer ex Forster 1846:373 pro syn.
      Cereus leptacanthus De Candolle ex Salm-Dyck 1845:27.
      Echinocereus leptacanthus Gaillard 1864:209.
      Echinocereus pentalophus var. subarticulatus* Weniger 1970:51 nom. inval.
Echinocereus pentalophus var. subarticulatus* Weniger 1970:51 nom. inval. (no basionym given).
  = Cereus pentalophus var. subarticulatus.
*Echinocereus pentlandii = Echinopsis pentlandii.
Echinocereus perbellus Britton & Rose 1922:24-25.
  Type - locality: Texas, Big Springs; Rose & Standley 12215, 23 Feb.1910 (US 635015, 635016).
    Nomenclatural synonyms:
      Echinocereus reichenbachii var. perbellus (Britton & Rose) Benson 1969d:127.
      Echinocereus caespitosus var. perbellus* Weniger 1970:23 nom. inval.
      Echinocereus reichenbachii subsp. perbellus* Blum 1997 Seed List nom. nud.
        Note: Considered a taxonomic synonym of  E. reichenbachii subsp. caespitosus by Blum et al. (1998b:334).
*Echinocereus persolutus [from Peru and thus not an Echinocereus].
Echinocereus phaeacanthus* Cactus & Succulent Journal (U.S.) 1951:104 nom. nud.
Echinocereus phoeniceus* Lemaire 1868:57 nom. nud.
  [Probably = Echinocereus coccineus].
Echinocereus phoeniceus* Engelmann ex Rumpler 1886:788 nom. illegit.
  basionym: Cereus phoeniceus Engelmann 1856:284 which was a new name for Echinocereus coccineus Engelman 1848:94.
Echinocereus phoeniceus* var. albispinus Hort. ex Rumpler 1886:789.
Echinocereus phoeniceus* fa. brevispinus Engelmann ex Schelle 1907:138 nom. nud.
Cereus phoeniceus subsp. conoideus Engelmann 1857:37.
  = Echinocereus conoideus.
Echinocereus phoeniceus* var. conoideus Schumann 1896:150.
  = Echinocereus conoideus.
Echinocereus phoeniceus* fa. densus Schelle 1907:138 nom. nud.
Echinocereus phoeniceus* var. inermis Schumann 1896:150.
  (see Echinocereus coccineus var. inermis).
Echinocereus phoeniceus* var. longispinus Hort. ex Rumpler 1886:789.
Cereus phoeniceus var. pacificus Engelmann ex Orcutt 1886:46.
  [see Echinocereus pacificus].
Echinocereus phoeniceus* var. rufispinus Hort. ex Rumpler 1886:789.
Echinocereus phoeniceus* fa. utahensis* Schelle 1907:138 nom. nud.
Echinocereus pleiogonus Croucher 1878:290.
  Type - locality: unknown [based on the illustration perhaps an Echinopsis].
Echinocereus pleiogonus* Labouret ex Rumpler 1886:828-829 nom. illegit. (later homonym).
  basionym: Cereus pleiogonus Labouret 1853:317.
    Type - locality: unknown; ? (?).
Echinocereus plomosus* Kohres Catalog 1984. nom. nud.
  [= Lau 768 from Plomosos, Coahuila ].
Echinocereus polichephalus* Daul 1890:78 nom. nud.
Echinocereus polycephalus* Hort. Berol. ex Haage 1897 nom. nud.
Echinocereus polyacanthus Engelmann 1848a:104.
  Type - locality:Mexico, Chihuahua, Cosihuiriachi; Wislizenus, 1846 (MO 2016721).
    Nomenclatural synonyms:
      Cereus polyacanthus (Engelmann) Engelmann 1849:50.
      Echinocereus triglochidiatus var. polyacanthus (Engelmann) Benson 1944:253.
Echinocereus polyacanthus var. acifer* Parker 1996:4 nom. nud.
  = Echinocereus acifer.
Echinocereus polyacanthus subsp. acifer (Otto ex Salm-Dyck) Taylor 1997:9.
  = Echinocereus acifer.
Echinocereus polyacanthus var. acifer fa. huitcholensis* Parker 1996:4. nom. nud.
  = Echinocereus huitcholensis.
Echinocereus polyacanthus var. acifer fa. matthesianus* Parker 1996:4 nom. nud.
  = Echinocereus matthesianus*.
Echinocereus polyacanthus var. albispinus Hort. ex Schelle 1926:179.
  Type - locality: unknown; ? (?).
Echinocereus polyacanthus var. bergeanus Fobe 1911:56.
  Type - locality: unknown; ? (?).
Echinocereus polyacanthus var. bergeanus* Hort ex Schelle 1926:179 nom. illegit. (later homonym).
Echinocereus polyacanthus var. densus (Regel) Taylor 1984:159.
  = Echinopsis valida var. densa Regel 1852:295.
    Note: This name is used by Taylor (1984) as the name of Echinocereus acifer.
Echinocereus polyacanthus var. durangensis* Parker 1996:4 nom. nud.
  = Echinocereus durangensis.
Echinocereus polyacanthus var. durangensis fa. marksianus* Parker 1996:4 nom. nud.
  = Echinocereus marksianus.
Echinocereus polyacanthus var. galtieri (Rebut ex Fobe) Schelle 1926:179.
  = Echinocereus galtieri.
Echinocereus polyacanthus var. huitcholensis (Weber) Taylor 1988:82.
  = Echinocereus huitcholensis.
Echinocereus polyacanthus subsp. huitcholensis (Weber) Taylor 1997:9.
  = Echinocereus huitcholensis.
Echinocereus polyacanthus var. leeanus* Parker 1996:4 nom. nud.
  = Echinocereus leeanus.
Echinocereus polyacanthus var. longispinus Hort. ex Schelle 1926:179.
  Type - locality: unknown; ? (?).
Echinocereus polyacanthus var. mombergerianus* Parker 1996:4 nom. nud.
  = Echinocereus mombergerianus.
Echinocereus polyacanthus subsp. mombergerianus (Frank) Breckwoldt & Matylewicz 1996:15.
  = Echinocereus mombergerianus.
Echinocereus polyacanthus var. neomexicanus* Weniger 1970:44 nom. inval. (no basionym given).
  = Echinocereus neomexicanus.
Echinocereus polyacanthus var. nigrispinus Fobe 1911:55.
  Type - locality: unknown; ? (?).
Echinocereus polyacanthus var. nigrispinus* Hort. ex Schelle 1926:179 nom. illegit. (later homonym).
Echinocereus polyacanthus var. pacificus (Engelmann ex Orcutt) Taylor 1984:160.
  = Echinocereus pacificus.
Echinocereus polyacanthus subsp. pacificus (Engelmann ex Orcutt) Breckwoldt in Breckwoldt & Matylewicz 1996:15.
  = Echinocereus pacificus.
Echinocereus polyacanthus var. rosei* Weniger 1970:43 nom. inval. (no basionym given).
  = Echinocereus rosei.
Echinocereus polyacanthus var. rufispinus Fobe 1911:55-56.
  Type - locality: unknown; ? (?).
Echinocereus polyacanthus var. rufispinus* Hort. ex Schelle 1926:179 nom. illegit. (later homonym).
Echinocereus polycephalus* Blanc 1888:39 nom. nud.
Echinocereus polycephalus* Hort. Berol. ex Haage 1900:160 nom. nud.
Echinocereus polycephalus* var. albispinus* Frič 1924:121 nom. nud.
Echinocereus polycephalus* var. brevinigrispinus* Frič 1924:121 nom. nud.
Echinocereus polycephalus* var. nigrispinus* Frič 1924:121 nom. nud.
Echinocereus polycephalus* var. spinosissimus* Frič 1924:121 nom. nud.
Echinocereus polycephalus var. xphoeniceus* Frič 1924:121 nom. nud.
Echinocereus polypus* Frič 1924:121 nom. nud.
Echinocereus poselgeri Lemaire 1868:57 [A new name needed because of earlier homonym].
  = Cereus tuberosus Poselger 1853:131 nom. illegit. (later homonym) [not Cereus tuberosus Pfeiffer 1837.]
    Type - locality: Texas, near Rio Grande River, above Belleville; Poselger, 1850 (MO 2015481).
      Nomenclatural synonyms:
        Echinocereus tuberosus* Rumpler 1886: 131 nom. illegit.
        Cereus poselgeri Coulter 1896:398.
        Wilcoxia poselgeri (Lemaire) Britton & Rose 1909:434.
        Wilcoxia tuberosa* Berger 1929:134 nom. illegit.
Echinocereus poselgeri fa. australis* Parker 1996:6 nom. nud.
Echinocereus poselgeri subsp. gerherdii Waldeis et al. 2008:53.
  Type - locality: Mexico,Nuevo Leon, Municipio Garcia, Sierra el Fraile; Köhres 8964 (KR 20011).
Echinocereus poselgeri subsp. kroenleinii (Cartier) Lange 1995:138.
  = Wilcoxia kroenleinii Cartier 1980 (Succulentes 2(2) :2-3). [see Echinocereus kroenleinii].
Echinocereus poselgeri fa. kroenleinii* Parker 1996:6 nom. nud.
  = Echinocereus kroenleinii.
Echinocereus poselgeri fa. tamaulipensis* Parker 1996:6 nom. nud.
  = Echinocereus tamaulipensis.
Echinocereus poselgeri fa. tuberosa* Parker 1996:6 nom. nud.
Echinocereus poselgeri var. waldeisii* Parker 1996:6 nom. nud.
  = Echinocereus waldeisii.
Echinocereus poselgeri var. waldeisii fa. deherdtii* Parker 1996:6 nom. nud.
  = Echinocereus tamaulipesis var. deherdtii.
Echinocereus poselgerianus Linke 1857:239.
  Type - locality: Mexico; ? (?).
    Nomenclatural synonyms:
      Cereus poselgerianus (Linke) Berger 1905:80.
      Echinocereus blanckii var. poselgerianus (Linke) Backeberg 1960:1999.
        Note: usually considered a taxonomic synonym of Echinocereus berlandieri.
Echinocereus primolanatus* Schwarz ex Backeberg 1960:2043 nom. inval. (no type designated).
Echinocereus primolanatus Taylor 1985b:130.
  Type - locality: Mexico, Coahuila, Sierra Paila, north of Estacion Marte, cultivated by D. Parker, 1985 (K).
    Nomenclatural synonyms:
      Echinocereus primolanatus* Schwarz ex Backeberg 1960:2043 nom. inval. (no type designated).
Echinocereus princeps Foster 1861:163.
  Type - locality: unknown [Of unknown origin and perhaps from South America in which case it would not be an Echinocereus].
Echinocereus procumbens* Lemaire 1868:56 nom. nud.
  [Probably = Echinocereus procumbens Engelmann ex Rumpler].
Echinocereus procumbens (Engelmann) Engelmann ex Rumpler 1886:781.
  basionym: Cereus procumbens Engelmann 1849:50.
    Type - locality: Mexico, Tamaulipas, Burita at mouth of Rio Grande, St. Louis Volunteers, 1846 (MO).
      Nomenclatural synonyms:
        Echinocereus pentalophus var. procumbens (Engelmann) Fournier 1935:25.
        Echinocereus pentalophus fa. procumbens* Parker 1996:4 nom. nud.
        Echinocereus pentalophus subsp. procumbens (Engelmann) Blum & Lange in Blum et al.1998:129.
Echinocereus procumbens var. gracilior Dautw. ex Schelle 1926:165
  Type - locality: unknown; ? (?).
Echinocereus procumbens var. longispinus Hort. ex Hirscht 1902:135.
  Type - locality: unknown; ? (?).
Cereus propinquus Salm Dyck ex Otto 1833:366.
  = Echinocereus pentalophus var. simplex.
Echinocereus propinquus De Candolle ex Haage 1892:118.
  = Echinocereus pentalophus var. simplex.
Cereus propinquus var. subarticulatus* Pfeiffer ex Forster 1846:373 pro syn.
  = Echinocereus pentalophus var. subarticulatus.
Echinocereus pseudohempelii* Kunzler Cat. nom. nud.
  = Echinocereus kuenzleri.
Echinocereus pseudopectinatus (Taylor) Taylor 1989:74.
  = Echinocereus bristolii var. pseudopectinatus Taylor 1985b:120.
    Type - locality: Mexico, Sonora, Moctezuma; Lau 607, 1973, 21 June 1984 (K) , cultivated by Taylor.
      Nomenclatural synonyms:
        Echinocereus scopulorum subsp. pseudopectinatus* Blum & Lange in Blum et al. 1998b:342.
Echinocereus pubellus* Cactus and Succulents Journal (US) 7:5 1936 nom. nud.
Echinopsis pulchella var. flore kermesina* Haage ex Forster 1846:364. pro syn.
  = Echinocereus amoenus.
Echinopsis pulchella var. rosea* Labouret 1853:292. pro syn.
  = Echinocereus amoenus.
Echinocereus pulchellus (Martius) F. Seitz 1870:11. [see Taylor 1989:75 ].
  = Echinocactus pulchellus Martius 1832:342.
    Type - locality: Mexico, Hidalgo, Pachuca; Karwinsky (?).
      Nomenclatural synonyms:
        Cereus pulchellus Pfeiffer 1837:74.
        Echinonyctanthus pulchellus (Martius) Lemaire 1839:85.
        Echinopsis pulchella (Martius) Zuccarini ex Forster 1846:363.
          Note: Usually listed as Echinocereus pulchellus Schumann 1894:185.
Echinocereus pulchellus var. acanthosetus Arias & Guzman in Arias Montes et al. 1997:63.
  Type - locality: Mexico, Oaxaca, between Magdalena Jicotlan & Tepelmeme de Morelos; Arias, Guzman-Cruz & Gama 958,
    29 Oct. 1991 (MEXU).
  Nomenclatural synonyms:
    Echinocereus pulchellus subsp. acanthosetus (Arias & Guzman) Blum in Blum et al. 1998:261.
Echinocereus pulchellus subsp. acanthosetus (Arias & Guzman) Blum in Blum et al. 1998b:261.
  = Echinocereus pulchellus var. acanthosetus.
Echinocereus pulchellus var. amoenus (Dietrich) Forster ex Schumann 1897:253.
  = Echinocereus amoena.
Echinocereus pulchellus subsp. amoenus* Blum 1997 Seed List nom. nud.
  = Echinocereus amoenus.
Echinocereus pulchellus var. costatus Halda, Kupčak & Sladk.2000 in Acta Mus. Richnov. Sect. Nat., 7(1) : 34 (2000).
  Type - locality: San Luis Potosi/Zacatecas border; J.J. Halda; J. Sladkovský 8504131 (PR).
Echinocereus pulchellus var. sharpii Taylor 1989:75-77.
  Type - locality: Mexico, Nuevo Leon, near San Roberto; 1900m, 24o50´N 100 25´W, Gonzalez G. 1985 (seed), cult. N. P. Taylor,
    4 May 1988 (K).
  Nomenclatural synonyms:
    Echinocereus pulchellus subsp. sharpii (Taylor) Taylor 1997:9.
    Echinocereus sharpii* S & K Breckwoldt 1998:1. nom. nud.
    Echinocereus pulchellus subsp. sharpii.
Echinocereus pulchellus subsp. sharpii (Taylor) Taylor 1997:9.
  = Echinocereus pulchellus var. sharpii.
Echinocereus pulchellus subsp. sladkovskyi Halda & Kupčak 2000.Acta Mus. Richnov. Sect. Nat., 7(1): 34 (2000).
  Type - locality: Mexico, Hidalgo ; J.J. Halda; J. Sladkovský 850418033 (PR).
Echinocereus pulchellus subsp. venustus Blum & Rischer in Blum et al. 1998b:255.
    Type - locality: Mexico San Luis Potosi [Lau 1036]; W. Rischer, 5 Dec. 1974 (ZSS A 17984).
Echinocereus pulchellus var. weinbergii (Weingart) Taylor 1985a:272.
  = Echinocereus weinbergii.
Echinocereus pulchellus subsp. weinbergii (Weingart) Taylor 1997:9
  = Echinocereus weinbergii.
Echinocereus purpureus Lahman 1935:141-143.
  Type - locality: Oklahoma, near Medicine Park; Lahman (MO).
    Nomenclatural synonyms:
      Echinocereus caespitosus var. purpureus* Weniger 1970:23 nom. inval.
      Echinocereus reichenbachii var. purpureus* Parker 1996:5 nom. nud.
        Note: Considered a taxonomic synonym of E. reichenbachii subsp. caespitosus by Blum et al. (1998b:322).
Echinocereus radians Engelmann 1848a:104.
  Type - locality:Mexico, Chihuahua, Cosihuiriachi; Wislizenus, 1846 (MO).
    Nomenclatural synonyms:
      Cereus adustus var. radians (Engelmann) Engelmann 1849:50.
      Echinocereus adustus var. schwarzii fa. radians* Parker 1996:6 nom. nud.
        Note: Usually considered a taxonomic synonym of Echinocereus adustus.
Echinocereus raphicephalus Forster 1861:164.
  Type - locality: unknown; ? (?). [May be of South American origin and not an Echinocereus].
Echinocereus rayonesensis Taylor 1988:75-76.
  Type - locality: Mexico, Nuevo Leon, Rayones, Lau 1101 (K).
Echinocereus rayonesensis fa. barkeanus* Parker 1996:3 nom. nud.
  [Probably = Echinocereus barkeanus*].
Echinocereus rectispinus Peebles 1938:675.
  Type - locality: Arizona, hills near Nogales, 3900 ft.; Peebles SF 905 (US 1729266).
    Nomenclatural synonyms:
      Echinocereus fendleri var. rectispinus (Peebles) Benson 1944:259.
      Echinocereus fasciculatus var. rectispinus* Benson 1969d:131 pro syn.
      Echinocereus fendleri subsp. rectispinus (Peebles) Taylor 1997:9.
Echinocereus rectispinus var. robustus Peebles 1938:675.
  [see Echinocereus robustus Peebles].
Echinocactus reichenbachianus* Tersch. ex  Salm-Dyck 1842:20.
Echinocereus reichenbachianus* Linke 1857:83 nom. nud.
Echinocereus reichenbachianus Hort. ex Haage 1859:20. [See Echinocereus reichenbachii].
  = Echinocactus reichenbachianus Terscheck ex Fennel 1843:282 (Terscheck, Verzeichnis 3. According to Schumann 1897:271).
    Type - locality: Mexico; ? (?).
      Nomenclatural synonyms:
        Echinopsis pectinata var. reichenbachiana (Terscheck) Salm-Dyck 1845:26.
        Cereus reichenbachianus (Terscheck) Labouret 1853:318.
Echinocereus reichenbachianus* Engelmann ex Haage 1893:23.
  Note: See note under Echinocereus reichenbachii.
Echinocereus reichenbachianus* Engelmann ex Seitz 1878:11 nom. nud.
Echinocereus reichenbachianus* Engelmann ex Haage 1893:23 (according to Index Kewensis 2:813) nom. illegit. (later homonym).
Cereus reichenbachianus var. castaneus (Engelmann) Labouret 1853:319.
  = Echinocereus caespitosus var. castaneus.

Echinocereus reichenbachii – Echinocereus zuniensis

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Echinocereus reichenbachii (Terscheck ex Walpers) Haage Jr. ex Britton & Rose 1922:25.
  = Echinocactus reichenbachii Terscheck ex Walpers 1843:320 (Terscheck Verzeichnis 2. According to Schumann 1897:452).
    Type - locality: Mexico.; ? (?).
      Nomenclatural synonyms:
        Echinocereus pectinatus var. reichenbachii (Terscheck ex Walpers) Werdermann 1930:t5.
        Echinocereus caespitosus var. reichenbachii* Ters. ex Borg 1937:174 nom. illegit.
          Note: Since Terscheck's Original publication has not been found there is confusion concerning whether the two names
            Echinocactus reichenbachii and Echinocactus reichenbachianus refer to the same plant. The name Echinocereus
            reichenbachii is usually applied to Echinocereus caespitosus although this is uncertain since the original description of
            Echinocactus reichenbachii did not have a description of the plant's flower, and no collector or exact type locality is listed.
            Benson (1982:946) designated Palmer 511, near Saltillo, Mexico, 10 April 1905 (US 570013) as the neotype. Unless
            someone is able tolocate a copy of Terscheck's original publication questions will remain concerning the proper use of this
            name.
Echinocereus reichenbachii var. albertii Benson 1969d:127.
  Type - locality: Texas, Jim Wells Co., near Alice; Benson 16,550 (POM 317080).
    Nomenclatural synonyms:
      Echinocereus reichenbachii var. fitchii fa. albertii* Parker 1996:5 nom. nud.
      Echinocereus fitchii var. albertii* Mesa Garden Seed Catalog1996 nom. nud.
      Echinocereus fitchii subsp. alberti (Benson) Blum & Lange in Blum et al. 1998a:5. [see Blum et al. 1998b:313].
      Echinocereus reichenbachii subsp. albertii (L.D.Benson) Pilbeam 2011:117.
        Note: Echinocereus melanocentrus* Lowry 1936:20 is usually considered a synonym of this variety.
Echinocereus reichenbachii subsp. albertii (L.D.Benson) Pilbeam 2011:117.
  = Echinocereus reichenbachii var. albertii Benson 1969d:127.
Echinocereus reichenbachii var. albiflorus* Seela 1940:92 nom. nud.
Echinocereus reichenbachii var. albispinus (Lahman) Benson1969d:127.
  = Echinocereus albispinus Lahman 1935:143.
    Note: Benson uses this name instead of Echinocereus baileyi.
Echinocereus reichenbachii var. anigosanthus* Ito 1981. nom. nud.
Echinocereus reichenbachii var. armatus (Poselger) Taylor 1985b:133.
  = Echinocereus pectinatus var. armatus.
Echinocereus reichenbachii subsp. armatus (Poselger) Taylor 1997:9
Echinocereus pectinatus var. armatus.
Echinocereus reichenbachii var. aureiflorus* Seela 1940:92 nom. nud.
Echinocereus reichenbachii var. baileyi (Rose) Taylor 1985b:133.
  = Echinocereus baileyi.
Echinocereus reichenbachii subsp. baileyi (Rose) Taylor 1997:9.
  = Echinocereus baileyi.
Echinocereus reichenbachii var. baileyi fa. albispinus* Mesa Garden Seed Catalog 1996 nom. nud.
  = Echinocereus albuspinus.
Echinocereus reichenbachii subsp. burrensis Frank, 2000:77.
  Type - locality: Serranias del Burro, Coahuila, MX; G. R. W. Frank 042 (ZSS).
Echinocereus reichenbachii var. caespitosus* Mesa Garden Seed Catalog 1996 nom. nud.
Echinocereus reichenbachii fa. caespitosus* Parker 1996:5 nom. nud.
  = Echinocereus caespitosus.
Echinocereus reichenbachii subsp. caespitosus Blum & Lange in Blum et al.1998:324.
  = Echinocereus caespitosus.
Echinocereus reichenbachii var. chisoensis (Marshall) Benson 1969d:127.
  = Echinocereus chisoensis.
Echinocereus reichenbachii subsp. comanchensis Felix , Oldach & J. Oldach in  Blum et al. 2005:209 (-222; figs.).
  Type - locality: Comanche County, north of Lake Lawtonka, T. & J. Oldach 0229, 15 June 1990 (ZSS AX24462).
Echinocereus reichenbachii var. fitchii (Britton & Rose) Benson 1969d:127.
  = Echinocereus fitchii.
Echinocereus reichenbachii subsp. fitchii (Britton & Rose) Taylor 1997:9.
  = Echinocereus fitchii.
Echinocereus reichenbachii var. fitchii fa. albertii* Parker 1996:5 nom. nud.
  = Echinocereus reichenbachii var. albertii.
Echinocereus reichenbachii var. fitchii fa. melanocentrus* Parker 1996:5 nom. nud.
  = Echinocereus melanocentrus*.
Echinocereus reichenbachii fa. pailanus* Parker 1996:5 nom. nud.
Echinocereus reichenbachii var. perbellus (Britton & Rose) Benson 1969d:127.
  = Echinocereus perbellus.
Echinocereus reichenbachii subsp. perbellus* Blum 1997 Seed List nom. nud.
  = Echinocereus perbellus.
Echinocereus reichenbachii var. purpureus* Parker 1996:5 nom. nud.
  = Echinocereus purpureus.
*Echinocereus reichii (from Chile and therefor not an Echinocereus).
Echinocereus relictus B. Wellard, sp. nov. (Figs. 3, 4).
  Type - UTAH: “Limestone Road” S of Sun River Community, 1.25 mi N
  of Utah/Arizona border, 1 mi W of Port of Entry, 795 m, 37.018198°, -113.632523°, 14 Apr 2016, B. Wellard 578 with K. Wellard (holotype:
  UT; isotype: ASU).
  = Echinocereus engelmannii var. purpureus L.D. Benson
Echinocereus rigidispinus* Engelmann ex Haage 1859:20 nom. nud.
Echinocereus rigidissimus Hort. ex F. A. Haage 1897:13.
  = Echinocereus pectinatus var. rigidissimus.
Echinocereus rigidissimus* Rose 1909a:293 nom. illegit.(later homonym).
  = Echinocereus pectinatus var. rigidissimus.
Echinocereus rigidissimus var. rubispinus (Frank & Lau) Taylor 1984:175.
  = Echinocereus pectinatus var. rubispinus Frank & Lau 1982:32-35.
Echinocereus rigidissimus subsp. rubispinus (Frank & Lau) Taylor 1997:9.
  = Echinocereus pectinatus var. rubispinus.
Echinocereus rischeri (R.C.Romer) W.Rischer 2007:118
  = Echinocereus scheeri subsp. rischeri R.C.Romer 2004:8.
Echinocereus ritteri* Haage 1958:182 nom. nud. [A form of Echinocereus pentalophus]
Echinocereus robustior* Britton & Rose 1922:28 pro syn.
  = Echinocereus pectinatus var. rigidissimus.
Echinocereus robustus (Peebles) Peebles 1940:219.
  basionym: Echinocereus rectispinus var. robustus Peebles 1938:675.
    Type - locality: Arizona, Pima Co. Between Tucson & Sabino Canyon; Peebles SF896, 27 Aug 1935 (US 1729267).
      Nomenclatural synonyms:
        Echinocereus fendleri var. robustus* Benson 1944:259 nom. illegit. (later homonym).
        Echinocereus fasciculatus fa. robustus* Parker 1996:2 nom. nud.
          Note: This plant is usually treated as a taxonomic synonym of Echinocereus fasciculatus.
Echinocereus roemeri* Lemaire 1868:56 nom. nud.
Echinocereus roemeri (Engelmann) Engelmann ex Haage 1859:20.
  basionym: Cereus roemeri Engelmann 1849:50. [not Muhlenpfordt].
    Type - locality: Texas, granitic region of the Llano. Lindheimer, 1847 (MO).
      Nomenclatural synonyms:
        Echinocereus coccineus fa. roemeri* Parker 1996:4 nom. nud.
        Echinocereus roemeri Engelmann ex Rumpler 1886:792.
Echinocereus roemeri* (Muhlenpfordt) Rydberg 1906:146 nom. illeg.
  basionym: Cereus roemeri Muhlenpfordt 1848:19. [not Engelmann].
    Type - locality: Northern Texas; Roemer (?).
      Nomenclatural synonyms:
        Echinocereus coccineus subsp. roemeri Blum, Lange & Rutow in Blum et al.1998:433.
Echinocereus roetteri (Engelmann) Engelmann ex Rumpler 1886:829-830.
  basionym: Cereus roetteri Engelmann 1856:345. [see Engelmann 1859:33, t. 41, fig. 3-5].
    Type - locality: Texas, stony hills near Frontera [El Paso]; Wright, 1851 (MO 2016865).
      Nomenclatural synonyms:
        Cereus dasyacanthus var. minor* Engelmann 1856:278.
        Echinocereus pectinatus var. minor (Engelmann) Benson 1968:125.
        Echinocereus dasyacanthus fa. roetteri* Parker 1996:2 nom. nud.
          Note: This plant is probably a natural hybrid.
Echinocereus roetteri var. lloydii (Britton & Rose) Backeberg 1960:2027.
  = Echinocereus lloydii.
Echinocereus roetteri var. neomexicanus ( J.M.Coult. ) A.D.Zimmerman in Powell & Weedin 2004:211 pro sp. hybr.
  = Echinocereus pectinatus var. neomexicanus (J.M.Coult.) L.D.Benson 1944.
Echinocereus roseanus* Backeberg 1937:21 nom. nud.
Echinocereus roseanus* Backeberg 1960:2071 pro syn.
  = Echinocereus rosei.
Echinocereus rosei Wooten & Standley 1915:457.
  Type - locality: New Mexico, Dona Ana Co., hills near Agricultural College [Las Cruces]; Standley 1235, 1907 (US 535093).
    Nomenclatural synonyms:
      Echinocereus triglochidiatus var. rosei (Wooten & Standley) Marshall 1950:63.
      Echinocereus roseanus* Backeberg 1960:2071 pro syn.
      Echinocereus polyacanthus var. rosei* Weniger 1970:44 nom. inval.
      Echinocereus coccineus fa. rosei* Blum 1990:13 nom. inval.
      Echinocereus triglochidiatus var. neomexicanus fa. rosei* Blum 1990:12. pro syn.
      Echinocereus coccineus var. neomexicanus fa. rosei* Parker 1996:4 nom. nud.
      Echinocereus coccineus subsp. rosei Blum & Rutow in Blum et al. 1998b:426.
      Echinocereus coccineus var. rosei Zimmerman ex Powell & Weedin 2004:203.
        Note: This plant is often listed under the name Echinocereus triglochidiatus var. neomexicanus the type of which is probably
          a natural hybrid.
Echinocereus rosei var. jarillensis* Kunzler Cat. nom. nud.
Echinocereus rotatus Linke 1858:85.
  Type - locality: South Texas; ? (?).
    Nomenclatural synonyms:
      Echinocereus pectinatus var. rotatus (Linke) Linke ex Haage 1892:118.
        Note: This name is usually consisdered a taxonomic synonym of Echinocereus caespitosus.
Echinocereus rubescens Dams 1905:92 -93.
  Type - locality: unknown; ? (?).
    Note: This species is often considered to be a taxonomic synonym of Echinocereus dasyacanthus.
Echinocereus rufispinus Engelmann 1848a:104-105.
  Type - locality:Mexico, Chihuahua, Cosihuiriachi; Wislizenus, 19 May 1848 (MO 2016892).
    Nomenclatural synonyms:
      Cereus rufispinus (Engelmann) Engelmann 1849:50.
      Cereus pectinatus fa. rufispinus (Engelmann) Voss 1894:378.
      Echinocereus adustus var. schwarzii fa. rufispina* Parker 1996:6 nom. nud.
        Note: Usually considered a taxonomic synonym of Echinocereus adustus.
Echinocereus rungei Schumann 1895:124. [This is a new name for Echinocereus texensis* Runge].
  Type - locality: Texas; Runge. (?).
    Nomenclatural synonyms:
      Echinocereus texensis* Runge 1894:61. nom. illegit.(later homonym).
        Note: The plant is usually considered a taxonomic synonym of Echinocereus papillosus.
Echinocereus runyonii Orcutt 1926:5.
  Type - locality: Texas, mouth of Rio Grande River, Orcutt 1048, 1924 (?).
    Note: This plant is probably the same as Echinocereus procumbens.
Echinocereus russanthus Weniger 1969:41-42.
  Type - locality: Texas, Brewster Co., Christmas Mtns.; Weniger 712, (UNM).
    Nomenclatural synonyms:
      Echinocereus chloranthus var. russanthus (Weniger) Lamb ex Rowley 1974:7.
      Echinocereus viridiflorus var. russanthus (Weniger) Leuck 1980:34.
        Note: Echinocereus chloranthus var. vulpis-cauda* is probably a synonym of this species
Echinocereus russanthus var. cowperi* Hort. [HK 1287, from Cooks Range, Luna County,New Mexico].
  = Echinocereus chloranthus subsp. rhyolithensis.
Echinocereus russanthus var. fiehnii Trocha 1997:65-69.
  Type - locality: Mexico, Chihuahua, Canon de Santa Clara, Sierra del Nido; Lau 1076 (HBG1066).
    Nomenclatural synonyms:
      Echinocereus russanthus var. finnii* Kohres Catalog 1981. nom. nud.
      Echinocereus finnii* Hort. ex Taylor 1985 101 pro syn.
      Echinocereus fiehnii* Lau 1992:74.
      Echinocereus russanthus fa. finnii* Parker 1996:3 nom. nud.
      Echinocereus russanthus subsp. fiehnii (Trocha) Blum & Lange in Blum et al.1998:212.
Echinocereus russanthus subsp. fiehnii (Trocha) Blum & Lange in Blum et al.1998:212.
  = Echinocereus russanthus var. fiehnii.
Echinocereus russanthus var. finnii* Kohres Catalog 1981. nom. nud.
  = Echinocereus russanthus var. fiehnii.
Echinocereus russanthus fa. finnii* Parker 1996:3 nom. nud.
  = Echinocereus russanthus var. fiehnii.
Echinocereus russanthus subsp. milleri* Blum 1997 Seed List nom. nud.
  = Echinocereus milleri.
Echinocereus russanthus var. neocapillus* Mesa Garden Seed Catalog 1996 nom. nud.
  = Echinocereus chloranthus var. neocapillus.
Echinocereus russanthus var. vulpis-cauda* Kohres Catalog 1981 nom. nud.
  = Echinocereus chloranthus var. vulpis-cauda*
Echinocereus russanthus fa. vulpis-cauda* Parker 1996:3 nom. nud.
  = Echinocereus chloranthus var. vulpis-cauda*
Echinocereus russanthus var. weedinii* Mesa Garden Seed Catalog 1996 nom. nud.
  = Echinocereus viridiflorus var. weedinii.
Echinocereus russanthus subsp. weedinii Leuck ex Blum & Lange in Blum et al. 1998b:214-216.
  Type - locality: Texas, Jeff Davis Co., top of Timber Mtn. , 6400 ft.; Leuck & Weedin 193, 13 May 1978 (OKL).
    Nomenclatural synonyms:
      Echinocereus chloranthus var. livermorensis* Kolle 1978:34 nom. inval. (not published in printed form).
      Echinocereus viridiflorus var. weedinii Leuck 1980:37-39 nom. inval.
      Echinocereus russanthus var. weedinii* Mesa Garden Seed Catalog 1996 nom. nud.
      Echinocereus chloranthus subsp. weedinii* Hochstatter Seed List 1997 nom. nud.
      Echinocereus viridiflorus var. weedinii ( Leuck ex Blum & Lange ) A.D. Zimmerman 2004: 270.
Echinocereus salm-dyckianus Scheer 1856:291.
  Type - locality: Mexico, Chihuahua; ? (?).
    Nomenclatural synonyms:
      Cereus salm-dyckianus (Scheer) Hemsley 1880:545.
      Echinocereus scheeri var. salm-dyckianus* Anett ex Parker 1996:4 nom. nud.
Echinocereus salm-dyckianus subsp. bacanorensis Rischer & Trocha 1998:3-10.
  Type - locality: Mexico, Sonora, near Bacanora; Trocha AX18239 (ZSS).
    Nomenclatural synonyms:
Echinocereus klapperi subsp. bacanorensis (Rischer) Fursch & Rischer 2003.
Echinocereus santaritensis subsp. bacanorensis (Rischer) Rischer & Felix 2006.
Echinocereus salm-dyckianus var. gracilior* Haage 1900:161 nom. nud.
Echinocereus salm-dyckianus var. gracilior Hort. ex Schelle 1926:163.
  Type - locality: unknown; ? (?).
Echinocereus salm-dyckianus var. noctiflorus Heid 1944:27.
  Type - locality: Mexico, Chihuahua, Cerocauhui; R. T. Craig (CA 305550).
Echinocereus salm-dyckianus subsp. obscuriensis Blum 1994:79-83.
  = Echinocereus scheeri var. obscuriensis.
Echinocereus salmianus Hort. ex Rumpler 1886:809.
  Type - locality: unknown; ? (?).
    Nomenclatural synonyms:
      Cereus salmianus (Hort. ex Rumpler) Weber in Bois 1894:279.
Echinocereus scheeri fa. salmianus* Parker 1996:4 nom. nud.
Echinocereus saltillensis* Mats. ex Haage 1900:161 nom. nud.
Echinocereus sanborgianus (Coulter) Schumann 1897:174.
  basionym: Cereus sanborgianus Coulter 1896:391.
    Type - locality: Mexico, Baja California, rocky table lands south of San Borja; Gabb 9, 1867 (MO).
      Nomenclatural synonyms:
        Echinocereus brandegeei fa. sanborgianus* Parker 1996:1 nom. nud.
          Note: Usually considered a taxonomic synonym of Echinocereus brandegeei.
Echinocereus sandersii Orcutt 1926:5.
  Type - locality: California, Black Canyon, Providence Mountains, 5000ft; Gibson & Sanders, 1925 (?).
    Note: Usually considered a synonym of taxonomic Echinocereus mojavensis.
Echinocereus sangre de christo Schumann 1897:278.
  Type - locality: Mexico, Mexico, near Pachuca; ; ? (?).
Echinocereus sanguineus Blanc 1890:58.
  Type - locality: unknown; ? (?).
    Note: Probably a form of Echinocereus viridiflorus.
Echinocereus sanpedroensis* Raudonat & Rischer 1995:88-97. nom. inval. (type contains two gatherings)
  Type - locality: Mexico, Sonora, Rancho San Pedro, Rischer, May 1995 (ZSS).
Echinocereus sanpedroensis Raudonat & Rischer 2002:54
  Type - locality:Mexico, Sonora, Rancho San Pedro; Rischer 253 (ZSS).
    Note: Echinocereus subterraneus* Backeberg 1960:2012 is sometimes listed as a synonym of this plant.
Echinocereus santaritensis Blum & Rutow in Blum et al. 1998:373-375.
  Type - locality: Arizona, Santa Rita Mountains; Rutow 52-56, 16 Apr. 1996 (UA).
Echinocereus santaritensis subsp. bacanorensis (Rischer) Rischer & Felix 2006:91.
  = Echinocereus salm-dyckianus subsp. bacanorensis Rischer & Trocha 1998:3-10.
Echinocereus sarissophorus Britton & Rose 1922:38-39.
  Type - locality: Mexico, Coahuila, near Saltillo; Palmer 100, Apr. 1898 (US).
    Nomenclatural synonyms:
      Echinocereus enneacanthus fa. sarissophorus* Parker 1996:2 nom. nud.
Echinocereus scheeri* Lemaire 1868:57 nom. nud.
  [Probably = Echinocereus scheeri Salm-Dyck ex Scheer].
Echinocereus scheeri (Salm-Dyck) Salm-Dyck ex Scheer 1856:291.
  basionym: Cereus scheeri Salm-Dyck 1850:190-191.
    Type - locality: Mexico, Chihuahua; Potts (?).
      Nomenclatural synonyms:
        Echinocereus schlini* Britton & Rose 1922:43 pro syn.
Echinocereus scheeri var. australis* Taylor 1988:80 pro syn.
  = Echinocereus scheeri var. koehresianus.
Echinocereus scheeri var. compactus* Hort. ex Haage 1897:13 nom. nud.
Echinocereus scheeri var. gentryi (Clover) Taylor 1984:154.
  = Echinocereus gentryi.
Echinocereus scheeri subsp. gentryi (Clover) Taylor 1997:9.
  = Echinocereus gentryi.
Echinocereus scheeri var. gentryi cultivar 'Cucumis' (Clover) Taylor 1984:155.
  = Echinocereus cucumis.
Echinocereus scheeri var. gentryi fa. cucumis* Parker 1996:4 nom. nud. = Echinocereus cucumis.
Echinocereus scheeri var. gracilior Hildmann ex Schelle 1926:162.
  Type - locality: unknown; ? (?).
Echinocereus scheeri var. koehresianus Frank 1988:186-189.
  Type - locality: Mexico, Durango, between El Carrizo & Loberas; Lau 1143 (ZSS).
    Nomenclatural synonyms:
      Echinocereus koehresianus* Hort. ex Taylor 1988:80 pro syn.
      Echinocereus koehresii* Hort. ex Taylor 1988:80 pro syn.
      Echinocereus scheeri var. australis* Taylor 1988:80 pro syn.
      Echinocereus ortegae subsp. koehresianus (Frank) Rischer & Frank 1996:93.
      Echinocereus ortegae fa. koehresianus* Parker 1996:4 nom. nud.
        Note: Taylor (1993) lists this species as a taxonomic synonym of Echinocereus ortegae.
Echinocereus scheeri var. major Rother 1905:175.
  Type - locality: unknown; ? (?).
Echinocereus scheeri var. minor Rother 1905:175.
  Type - locality: unknown; ? (?).
Echinocereus scheeri var. nigrispinus* Scheer 1856:291 nom. nud.
Echinocereus scheeri var. obscuriensis Lau 1989:34-36.
  Type - locality: Mexico, Chihuahua, Sierra Obscura; Lau 091 (ZSS).
    Nomenclatural synonyms:
      Echinocereus salm-dyckianus subsp. obscuriensis (Lau) Blum 1994:79.
Echinocereus scheeri var. salm-dyckianus fa. obscuriensis* Parker 1996:4 nom. nud.
Echinocereus scheeri var. obscuriensis* Lau in Guzman et al. 2003a:78 (without basionym).
Echinocereus scheeri subsp. obscuriensis (Lau) Guzman 2003b:17.
  = Echinocereus scheeri var. obscuriensis Lau published without basionym in Guzman et al. 2003a:78.
Echinocereus scheeri subsp. paridensis Rischer & Trocha 1998:58-65.
  Type - locality: Mexico , Sinaloa, Near Parida; Rischer 1992/312, 1992 (ZSS AX 18719).
Echinocereus scheeri subsp. polyacanthoides* Momberger 1994:58. nom. prov. (from Tomochic, Chihuahua, Mexico).
Echinocereus scheeri subsp. rischeri R.C.Romer 2004:8.
  Type - locality: Mexico, Chihuahua, Municipio Guachochi, Cumbres de Sinforosa; Werner Rischer WR 605 (UAT).
Echinocereus scheeri var. robustior Rother 1902:59.
  Type - locality: unknown; ? (?).
Echinocereus scheeri var. salm-dyckianus* Anett ex Parker 1996:4 nom. nud.
  = Echinocereus salm-dyckianus.
Echinocereus scheeri var. salm-dyckianus fa. obscuriensis* Parker 1996:4 nom. nud.
  = Echinocereus scheeri var. obscuriensis.
Echinocereus scheeri fa. salmianus* Parker 1996:4 nom. nud.
  = Echinocereus salmianus.
Echinocereus schereri Frank 1990a:154-159.
  Type - locality: Mexico, Durango,near Las Minas, Rio Nazas; Scherer 123 (ZSS).
Echinocereus schlechterdalii* Britton & Rose 1922:302 error for Echinocactus schlechtendalii.
Echinocereus schlini* Britton & Rose 1922:43 pro syn.
  = Echinocereus scheeri.
Echinocereus schmollii (Weingart) Taylor 1985b:140.
  basionym: Cereus schmollii Weingart 1931:251-252.
    Type - locality: Mexico, Queretero; F. Schmoll (?).
      Nomenclatural synonyms:
        Wilcoxia schmollii (Weingart) Backeberg 1935, Blatter fur Kakteenforschung. 2(11) :93.
          Note: Wilcoxia nerispina* Backeberg 1960:2078 nom. invalid. is usually considered a synonym of this species.
Wilcoxia schmollii var. lanata* Cartier 1980, Succulentes 2(4) :8 nom. inval.
Echinocereus schmollii fa. nerispina* Parker 1991:6 nom. nud.
  = Wilcoxia nerispina*
Echinocereus schwarzii* Backeberg 1960:2032 pro syn.
  = Echinocereus radians.
Echinocereus schwarzii Lau 1982:27-29.
  Type - locality: Mexico, Durango, near Guenacevi, 1800-2100 m.; Lau 1305 (POM).
    Nomenclatural synonyms:
      Echinocereus adustus var. schwarzii (Lau) Taylor 1985a:268.
      Echinocereus adustus subsp. schwarzii (Lau) Taylor 1992:8.
Echinocereus sciurus (K. Brandegee) K. Brandegee ex Dams 1904:130.
  basionym: Cereus sciurus K. Brandegee 1904:192.
    Type - locality: Mexico, Baja California Sur, hills near San Jose del Cabo; T.S. Brandegee, April 1897 (UC).
Echinocereus sciurus var. floresii (Backeberg) Taylor 1985b:115.
  = Echinocereus floresii.
Echinocereus sciurus subsp. floresii (Backeberg) Taylor 1997:10.
  = Echinocereus floresii.
Echinocereus sciurus var. floresii fa. townsendianus* Parker 1996:5 nom. nud.
  = Echinocereus townsendianus*
Echinocereus scopulorum Britton & Rose 1922:30-31.
  Type - locality: Mexico, Sonora, near Guaymas; Rose et al 12570, 10 March 1910 (US).
    Nomenclatural synonyms:
      Echinocereus pectinatus var. scopulorum (Britton & Rose) Marshall 1955:61.
Echinocereus scopulorum subsp. pseudopectinatus Blum & Lange in Blum et al. 1998b:342.
  = Echinocereus pseudopectinatus.
*Echinocereus serpentinus Lemaire= Nyctocereus serpentinus.
Echinocereus setosus* Hort. ex Taylor 1988:76 pro syn.
  = Echinocereus freudenbergeri.
Cereus similis* Regel & Klein 1860:29. according to Hemsley 1880:546, but not found on that page.
  Type - locality: Mexico; Karwinsky (?).
    Note: Perhaps this is the same as Cereus similis Meinshausen 1858 in Wochenschrift für Gärtnerei und Pflanzenkunde 1:29].
*Echinocereus spachianus Rumpler = Echinopsis spachiana.
*Echinocereus spinibarbis Salm Dyck = Eulychnia spinibarbis.
Echinocereus spinibarbis* Haage 1900:161 nom. nud.
Echinocereus spinibarbis* Hildmann ex Fobe 1911:58 nom. illegit. (later homonym).
Echinocereus spinigemmatus Lau 1984:248-250.
  Type - locality: Mexico, Jalisco, near San Andres Cohamiata, 1600 m.; Lau 1246, 25 Apr. 1974 (ZSS AA18- 48).
Echinocereus spinosissimus Walton 1899a:162.
  Type - locality: Texas, El Paso; ? (?).
    Note: Usually considered a taxonomic synonym of Echinocereus dasyacanthus.
Echinocereus spinosissimus var. ctenoides* (Engelmann) Ito 1981:237. nom. inval. (no basionym).
  = Echinocereus ctenoides.
Echinocereus spinosissimus var  steereae* (Clover) Ito 1981:237. nom. inval. (no basionym).
  = Echinocereus steereae Clover.
Echinocereus spinosus* Coulter ex Schelle 1926:174 pro syn.
  = Echinocereus pectinatus var. armatus.
*Echinocereus splendens Lemaire.
  = Nyctocereus splendens.
Echinocereus standleyi Britton & Rose 1922:24.
  Type - locality: New Mexico, Sacramento Mtns.,100 miles north of El Paso; S. L. Pattison (US).
    Nomenclatural synonyms:
      Echinocereus viridiflorus var. standleyi (Britton & Rose) Orcutt ex Weniger 1969:37.
      Echinocereus viridiflorus fa. standleyi* Parker 1996:3 nom. nud.
Echinocereus steereae Clover 1938:565-567.
  Type - locality: Texas, Chisos Mtns.; Mrs Lois Steere (MICH).
    Nomenclatural synonyms:
      Echinocereus dasyacanthus var. steereae (Clover) Marshall 1945:115.
      Echinocereus pectinatus var. steereae* Rumpler ex Weniger 1970:32 pro syn.
      Echinocereus spinosissimus var  steereae* (Clover) Ito 1981:237. nom. inval. (no basionym).
      Echinocereus dasyacanthus fa. steereae* Parker 1996:2 nom. nud.
Echinocereus stoloniferus Marshall 1938:159-160.
  Type - locality: Mexico, Sonora, Mountain facing Rancho Guirocoba, 5000 ft.; Bristol & Marshall, 19 Mar. 1935 (DS 251176).
    Nomenclatural synonyms:
      Echinocereus subterraneus* Hort. ex Bailey 1941:271. pro. syn.
Echinocereus stoloniferus var. tayopensis (Marshall) Taylor 1985a:258.
  = Echinocereus tayopensis.
Echinocereus stoloniferus subsp. tayopensis* (Marshall) Pichler 1994:103-106. nom. inval.
  = Echinocereus tayopensis.
Echinocereus stoloniferus subsp. tayopensis (Marshall) Taylor 1997:10.
  = Echinocereus tayopensis.
Echinocereus stramineus (Engelmann) Engelmann ex Seitz 1870:11.
  basionym: Cereus stramineus Engelmann 1856:282. [see Engelmann 1859:35, t. 46].
    Type - locality: Texas, Mtns near El Paso; Wright, 1851 (MO).
      Nomenclatural synonyms:
        Echinocereus enneacanthus var. stramineus (Engelmann) Benson 1969d:127.
          Note:Usually cited as Echinocereus stramineus Engelmann ex Rumpler 1886:797.
Echinocereus stramineus var. conglomeratus (Blanc) Bravo 1974:47
  = Echinocereus conglomeratus.
Echinocereus stramineus fa. conglomeratus* Parker 1996:3 nom. nud.
  = Echinocereus conglomeratus.
Echinocereus stramineus var. major* Haage 1897:13 nom. nud.
Echinocereus stramineus fa. major* Schelle 1907:136 nom. nud.
Echinocereus stramineus var. major-aureispina* Frič 1924:121 nom. nud.
Echinocereus stramineus var. occidentalis Taylor 1988:70-71.
  Type - locality: Mexico, Durango, southeast of La Cuesta, 140 km north of city of Durango, on volcanic rocks, 1400-1450 m.;
    N.P.Taylor 240b, 1 July 1986 (MEXU).
  Nomenclatural synonyms:
    Echinocereus stramineus subsp. occidentalis (Taylor) Taylor 1997:10.
    Echinocereus occidentalis (Taylor) Richer et al. 2009:23.
Echinocereus stramineus subsp. occidentalis (Taylor) Taylor 1997:10.
  = Echinocereus stramineus var. occidentalis.
Echinocereus stramineus var. parkeri* Seed list 1996 - British Cactus and Succulent Journal. = Echinocereus parkeri.
Echinocereus stramineus var. ruberrimus* Frič 1924:121 nom. nud.
Echinocereus stramineus var. spinarosa* Frič 1924:121 nom. nud.
Echinocereus strausianus Haage ex Quehl 1900:70-71.
  Type - locality: Texas; ? (?).
    Note: Usually considered a taxonomic synonym of Echinocereus viridiflorus var. cylindricus.
*Echinocereus strigosus Lemaire = Echinopsis strigosa.
Echinocereus subinermis Salm-Dyck ex Scheer 1856:291.
  Type - locality: Mexico, Chihuahua, Potts (?).
    Nomenclatural synonyms:
      Cereus subinermis (Salm-Dyck ex Scheer) Hemsley 1880:546.
Echinocereus subinermis var. aculeatus Unger 1984:164.
  Type - locality: Mexico, Chihuahua, Rio Batopilas near la Bufa; 700m, Unger, 1980 (ZSS AA50- 70).
    Nomenclatural synonyms:
      Echinocereus subinermis fa. aculeatus* Parker 1996:4 nom. nud.
Echinocereus subinermis fa. aculeatus* Parker 1996:4 nom. nud.
  = Echinocereus subinermis var. aculeatus.
Echinocereus subinermis var. luteus (Britton & Rose) Knuth in Backeberg & Knuth 1935:312.
  = Echinocereus luteus.
Echinocereus subinermis fa. luteus (Britton & Rose) Krainz 1967:51.
  = Echinocereus luteus.
Echinocereus subinermis var. ochoterenae Unger 1984:164.
  = Echinocereus ochoterenae.
Echinocereus subinermis subsp. ochoterenae (Ortega) Taylor 1997:10.
  = Echinocereus ochoterenae.
Echinocereus subterraneus* Hort. ex Bailey 1941:271 pro syn.
  = Echinocereus stoloniferus.
Echinocereus subterraneus* Backeberg 1937:21 nom. nud.
Echinocereus subterraneus* Backeberg 1960:2012 nom. inval. (no type designated).
  Type - locality: Mexico [See Echinocereus sanpedroensis].
Echinocereus tamaulipensis* Frič 1926:441 nom. nud.
  [probably = Echinocereus pectinatus var. tamaulipensis].
Echinocereus tamaulipensis (Werdermann) Lange 1995:139-140.
  = Wilcoxia tamaulipensis Werdermann 1938:86.
    Type - locality: Mexico, Southern Coahuila, Hummel s.n. (B).
      Nomenclatural synonyms:
        Echinocereus poselgeri fa. tamaulipensis* Parker 1996:6 nom. nud.
Echinocereus tamaulipensis subsp. deherdtii Lange 1995:140-141.
  Type - locality: Mexico, Coahuila, El Sol; Lange (JE).
    Nomenclatural synonyms:
      Echinocereus poselgeri var. waldeisii fa. deherdtii* Parker 1996:6 nom. nud.
Echinocereus tamaulipensis subsp. waldeisii (Haugg) Lange 1995:142.
  = Echinocereus waldeisii.
Echinocereus tayopensis Marshall 1956a:78-80.
  Type - locality: Mexico, Sonora, Rancho Saucito, between Bacadehuachi & Nacori Chico, 3500 ft (29o42'N 109o04'W); D. Gold & Sanchez-Mejorada (DES).
    Nomenclatural synonyms:
      Echinocereus stoloniferus var. tayopensis (Marshall) Taylor 1985a:258.
      Echinocereus stoloniferus subsp. tayopensis (Marshall) Pichler 1994:103-106.
Echinocereus texensis Jacobi 1856:110. [not Runge 1894].
  Type - locality: Texas.
    Note: This name is usually considered a taxonomic synonym of Echinocereus caespitosus.
Echinocereus texensis* Runge 1894:61-62 nom. illegit. (later homonym).
  [See Echinocereus rungei.]
Echinocereus thurberi* Monatschrift für Kakteenkunde 3:153. error for Cereus thurberi.
Echinocereus topiensis Rischer & Trocha 1999:9.
  Type - locality: Mexico, Durango, Topia; Anon 1992/292 (HBG 1118).
    Nomenclatural synonyms:
      Echinocereus acifer subsp. topiensis (W. Rischer & Trocha) W. Rischer 2004:88.
Echinocereus toroweapensis (Fischer) Fuersch 1993:266-268.
  = Echinocereus triglochidiatus var. toroweapensis.
Echinocereus townsendianus* Heyer 1989:26 nom. nud.
  [a form of Echinocereus floresii].
Echinocereus thwaitesii* Schumann 1897:290 nom. nud.
Echinocereus trichacanthus Rumpler 1886:799.
  Type - locality: unknown; ? (?).
*Echinocereus trichacanthus Engelmann ex Haage 1893:23.
  = Cephalocereus nobilis.
Echinocereus triglochidiatus Engelmann 1848a:93.
  Type - locality: New Mexico, Wolf Creek; Wislizenus 516, 24 June 1846 (MO 2016786).
    Nomenclatural synonyms:
      Cereus triglochidiatus (Engelmann) Engelmann 1849:50.
      Echinocereus paucispinus var. triglochidiatus* Schumann 1897:281 nom. illegit.
Echinocereus triglochidiatus var. acifer (Jacobi) Bravo 1978:66.
  = Echinocereus acifer.
Echinocereus triglochidiatus subsp. acifer (Otto ex Salm-Syck) Guzman 2003b:17.
  = Echinocereus acifer [published without basionym in Guzman et al. 2003a:78].
Echinocereus triglochidiatus var. arizonicus (Orcutt) Benson 1969a:21.
  = Echinocereus arizonicus.
Echinocereus triglochidiatus var. coccineus (Engelmann) Engelmann ex Marshall & Bock 1941:117.
Echinocereus triglochidiatus subsp. coccineus (Engelmann) Guzman 2003b:17.
  = Echinocereus coccineus. [published without basionym in Guzman et al. 2003a:78].
*Echinocereus triglochidiatus subsp. glochidiatus  Hunt et. al 2006:83{Typographical error]
  = Echinocereus triglochidiatus subsp. triglochidiatus
Echinocereus triglochidiatus var. gonacanthus (Engelmann & Bigelow) Boissevain 1940:36.
  = Echinocereus gonacanthus.
Echinocereus triglochidiatus var. gurneyi Benson 1969d:126.
  Type - locality: Texas, Brewster Co., igneous rocks south of Marathon; Correll & Benson 16,488 (POM).
    Nomenclatural synonyms:
      Echinocereus coccineus var. gurneyi (Benson) Heil & Brack1988:26.
      Echinocereus coccineus subsp. gurneyi * Blum 1997 Seed List nom. nud.
      Echinocereus coccineus subsp. gurneyi (L. Benson) D. Felix & H. Bauer 2014. - EcJ-Online 2 Sonderheft: XIII
        Note: This species is now believed to be a natural hybrid.
Echinocereus triglochidiatus var. hexaedrus (Engelmann & Bigelow) Boissevain 1940:36.
  = Echinocereus hexaedrus.
Echinocereus triglochidiatus var. huitcholensis (Weber) Guzman 2003b:17
  = Echinocereus huitcholensis [published without basionym in Guzman et al. 2003a:79].
Echinocereus triglochidiatus var. inermis Arp 1973:132-133
  = Echinocereus coccineus var. inermis.
Echinocereus triglochidiatus var. melanacanthus (Engelmann) Benson 1944:254.
  = Echinocereus coccineus var. melanacanthus.
    Note: This is the name used by Benson for Echinocereus coccineus.
Echinocereus triglochidiatus var. melanacanthus fa. inermis* Alabaster 1994:105 nom. inval.
  = Echinocereus coccineus var. inermis.
Echinocereus triglochidiatus var. mojavensis (Engelmann & Bigelow) Benson 1944:255.
  = Echinocereus mojavensis.
Echinocereus triglochidiatus subsp. mojavensis (Engelmann & Bigelow) Blum & Lange in Blum et al. 1998b:362.
  = Echinocereus mojavensis.
Echinocereus triglochidiatus var. mojavensis fa. inermis (Schumann) Ferguson 1989:219.
  = Echinocereus coccineus var. inermis.
Echinocereus triglochidiatus subsp. mojavensis fa. inermis (Schumann) Blum in Blum et al. 1998a:9. [see Blum et al. 1998b:365].
  = Echinocereus coccineus var. inermis.
Echinocereus triglochidiatus var. multicostatus (Cels ex Rumpler) Marshall 1953:67.
  = Echinocereus multicostatus Cels ex Rumpler 1886:834.
Echinocereus triglochidiatus var. neomexicanus (Standley) Standley ex Marshall & Bock 1941:118.
  = Echinocereus neomexicanus.
Echinocereus triglochidiatus var. neomexicanus fa. rosei* Blum 1990:12. pro syn.
  = Echinocereus rosei.
Echinocereus triglochidiatus var. octacanthus* Muhlenpfordt ex Marshall & Bock 1941:118 nom. invalid.
  = Echinocereus octacanthus.
Echinocereus triglochidiatus var. pacificus (Englemann ex Orcutt) Engelmann ex Marshall & Bock 1941:118.
  = Echinocereus pacificus.
Echinocereus triglochidiatus subsp. pacificus (Englemann ex Orcutt) Guzman 2003b:17.
  = Echinocereus pacificus [published without basionym in Guzman et al. 2003a:79.]
Echinocereus triglochidiatus var. paucispinus (Engelmann) Engelmann ex Marshall & Bock 1941:118.
  = Echinocereus paucispinus.
Echinocereus triglochidiatus var. polyacanthus (Engelmann) Benson 1944:253.
  = Echinocereus polyacanthus.
Echinocereus triglochidiatus subsp. polyacanthus (Engelmann) Guzman 2003b:17.
  = Echinocereus polyacanthus [published without basionym in Guzman et al. 2003a:79.]
Echinocereus triglochidiatus var. rosei (Wooten & Standley) Marshall 1950:63.
  = Echinocereus rosei.
Echinocereus triglochidiatus var. toroweapensis Fischer 1991:195.
  Type - locality: Arizona, Mohave Co.,Toroweap Point 1340 m.; P.C. Fischer 7196 (ARIZ).
    Nomenclatural synonyms:
      Echinocereus toroweapensis (Fischer) Fuersch 1993:266-268.
Echinocereus triglochititus* Engelm. ex Haage 1900:162. nom. nud.
  [probably an error in spelling for Echinocereus triglochidiatus].
Echinocereus trockyi* Frič 1918(8) [in advertisement] nom. nud.
Echinocereus tuberculatus* Hort. ex Haage 1900:162 nom. nud.
Echinocereus tuberosus* Poselger ex Rumpler 1886:783 nom. illegit.
  basionym: Cereus tuberosus Poselger 1853:135 non Pfeiffer.
  = Echinocereus poselgeri.
Echinocereus tulensis Bravo 1973:110-111.
  Type - locality: Mexico, Tamaulipas, 1 km. south of Tula Junction on highway between Huizache & Juamave;
    Sanchez-Mejorada 2085, 4 May 1973 (MEXU).
  Nomenclatural synonyms:
    Echinocereus cinerascens var. tulensis (Bravo) Taylor 1988:69.
    Echinocereus cinerascens subsp. tulensis (Bravo) Taylor 1997:8.
Echinocereus twaitesii* Hort. ex Haage 1892:119 nom. nud.
Echinocereus uehri Blanc 1888:40.
  Type - locality: unknown.
Echinocereus undulatus* Hildmann ex Schumann 1895:124 pro syn.
  = Echinocereus cinerascens.
Echinocereus uspenskii Blanc 1888:40.
  Type - locality: unknown; ? (?).
    Nomenclatural synonyms:
      Echinocereus enneacanthus fa. uspenskii* Parker 1996:2 nom. nud.
Echinocereus uspenskii* Haage 1900:162 nom. illegit. (later homonym).
Echinocereus utahensis* Kohres Cat 1984. nom. nud.
Echinocereus vatteri* Botzenhart Kakteen und andere Sukkulenten 1968 front cover nom. nud.
  = Echinocereus viereckii.
Echinocereus viereckii Werdermann 1934:188-189.
  Type - locality: Mexico, Tamaulipas, Mtns. between Juamave & Cd. Victoria, 1500-2000 m. ; ? (?).
Echinocereus viereckii subsp. huastecensis Blum, Lange & Rutow in Blum et al.1998:156-157.
  Type - locality: Mexico, Nuevo Leon, Rayones Canyon, 15 km west of  highway 85 on state road 6, 9 June 1986; Taylor 192 (K).
Echinocereus viereckii var. morricalii (Riha) Taylor 1985b:93.
  = Echinocereus morricalii.
Echinocereus viereckii subsp. morricalii (Riha) Taylor 1997:10.
  = Echinocereus morricalii.
Echinocereus viereckii subsp. santamariensis Blum & Felix 2010:65.
  Type - locality: Mexiko: Nuevo Leon: 5 km along road from Rayones to Galeana, 1000 m.; N.P. Taylor 194, 9 June 1986, [K].
Echinocereus virickii* White 1948:288 nom. nud.
Echinocereus viridiflorus Engelmann 1848a:91.
  Type - locality: New Mexico, Wolf Creek, Wislizenus 514, (MO 2016902).
    Nomenclatural synonyms:
      Cereus viridiflorus (Engelmann) Engelmann 1849:50.
      Echinocereus labouretii* Forster ex Rumpler 1886:811 pro syn.
Echinocereus viridiflorus var. canus Powell & Weedin 2004:273.
  Type - locality: Presidio Co., Solitario, S-facing crest of a novaculite ridge, c. 4847ft; A.M.Powell & S.A.Powell 6012 (SRSC).
    Nomenclatural synonym:
      Echinocereus canus (Powell & Weedin) Felix & Bauer in Blum et al. 2012:258 - 276.
Echinocereus viridiflorus var. centrispinus* Bailey 1941:271 nom. nud.
Echinocereus viridiflorus var. chloranthus (Engelmann) Backeberg 1960:2015.
  = Echinocereus chloranthus.
Echinocereus viridiflorus fa. chloranthus (Engelmann) Krainz 1967:51.
  = Echinocereus chloranthus.
Echinocereus viridiflorus subsp. chloranthus (Engelmann) Taylor 1997:10.
  = Echinocereus chloranthus.
Echinocereus viridiflorus var. chrysacanthus* Schelle1907:129 nom. nud.
Echinocereus viridiflorus var. correllii Benson 1969d:128.
  Type - locality: Texas, Brewster Co., above Ft. Pena Colorada, 4 mi. south of Marathon,4100 ft.; Benson & Correll 16485, 4 Apr.(POM317079).
    Nomenclatural synonyms:
      Echinocereus viridiflorus subsp. correllii (Benson) Blum & Lange in Blum et al. 1998b:198.
Echinocereus viridiflorus subsp. correllii (Benson) Blum & Lange in Blum et al. 1998b:198.
  =Echinocereus viridiflorus var. correllii.
Echinocereus viridiflorus var. cylindricus (Engelmann) Engelmann ex Rumpler 1886:812.
  basionym: Cereus viridiflorus var. cylindricus Engelmann 1856:278.
    Type - locality: Texas, Valley of Limpia Creek; Wright [418], 11 June 1851 (MO 2016904).
      Nomenclatural synonyms:
        Echinocereus chloranthus var. cylindricus (Engelmann) Taylor 1984:169.
        Echinocereus viridiflorus subsp. cylindricus (Engelmann) Taylor 1991:10.
        Echinocereus chloranthus subsp. cylindricus Blum & Lange in Blum et al. 1998b:205.
          Note: Echinocereus strausianus is usually considered a taxonomic synonym of this variety.
Echinocereus viridiflorus var. davisii (Houghton) Marshall & Bock 1941:119.
  = Echinocereus davisii.
Echinocereus viridiflorus fa. davisii (Houghton) Krainz 1967:51.
  = Echinocereus davisii.
Echinocereus viridiflorus subsp. davisii (Houghton) Taylor 1997:10.
  = Echinocereus davisii.
Echinocereus viridiflorus var. elongatus* Hort. ex Backeberg 1962:3847 nom. nud.
Echinocereus viridiflorus fa. faciliflorus* Hildmann ex Schelle 1907:129 nom. nud.
Echinocereus viridiflorus var. gracilispinus Hort. ex Rumpler 1886:814.
  Type - locality: unknown; ? (?).
Echinocereus viridiflorus var. intermedius* Backeberg 1960:2015 nom. inval. (no type designated).
  Note: This plant is said to be from Texas.
Echinocereus viridiflorus fa. longispinus* Schelle 1907:129 nom. nud.
Echinocereus viridiflorus var. major* Maas 1906:142 nom. nud.
Echinocereus viridiflorus var. minor* Engelmann ex Weniger 1970:14 nom. nud.
  basionym: Cereus viridiflorus var. minor* Engelmann 1856:278 which is a new name for Echinocereus viridiflorus var.
    viridiflorus.
Echinocereus viridiflorus var. montanus* Kunzler Catalog nom. nud.
Echinocereus viridiflorus var. neocapillus Leuck ex Zimmerman in Powell & Weedin 2004:267.
  = Echinocereus chloranthus var. neocapillus.
Echinocereus viridiflorus var. robustior* Kuenzler HK1007 Los Pinos, Valencia Co., NM
  Field number HK1007 Los Pinos, Valencia Co., NM
Echinocereus viridiflorus var. ruberrimus* Frič 1924:121 nom. nud.
Echinocereus viridiflorus var. rubrus* Frič 1924:121 nom. nud.
Echinocereus viridiflorus var. russanthus (Weniger) Zimmerman in Powell & Weedin 2004:263.
  = Echinocereus russanthus.
Echinocereus viridiflorus fa. sanguineus* Rgl. ex Schelle 1907:129 nom. nud. [perhaps the same as Echinocereus sanguineus].
Echinocereus viridiflorus var. standleyi (Britton & Rose) Orcutt ex Weniger 1969:37.
  = Echinocereus standleyi.
Echinocereus viridiflorus fa. standleyi*. Parker 1996:3 nom. nud.
  = Echinocereus standleyi.
Echinocereus viridiflorus var. tuberculosus* Orcutt 1926:4 nom. nud.
Echinocereus viridiflorus var. tubulosus* Heller 1900:8 nom. illegit.
  basionym: Cereus viridiflorus var. tubulosus* Coulter 1896:383.
    Note: new name for Echinocereus viridiforus var. cylindricus.
Echinocereus viridiflorus var. weedinii* Leuck 1980:37-39 nom. inval.
Echinocereus viridiflorus var. weedinii ( Leuck ex W.Blum & Mich.Lange ) A.D. Zimmerman 2004: 270.
  = Echinocereus russanthus subsp. weedinii.
Echinocereus waldeisii Haugg 1994:6.
  Type - locality: Mexico, San Luis Potosi, west of Tula, Huizache; D. Waldeis, 1988 (WU).
    Nomenclatural synonyms:
      Echinocereus tamaulipensis subsp. waldeisii (Haugg) Lange 1995:142.
      Echinocereus poselgeri var. waldeisii* Parker 1996:6 nom. nud.
Echinocereus webeserii* Pavey 1979: n.p. nom. nud.
  [Apparently a form of Echinocereus baileyi].
Echinocereus websterianus Lindsay 1947:153.
  Type - locality: Mexico, Gulf of California, San Pedro Nolasco Is.; Lindsay & Bool 498, 25 Feb. 1947 (DS314191).
Echinocereus weinbergii Weingart 1912:83.
  Type - locality: North America [This plant has been relocated near Sombrerete, Zacatecas, Mexico]; ? (?).
    Nomenclatural synonyms:
      Echinocereus pulchellus var. weinbergii (Weingart) Taylor 1985a:272.
      Echinocereus pulchellus subsp. weinbergii B(Weingart) Taylor 1997:9.
      Echinocereus weinbergii var. albispinus* Hort. ex Schelle 1926:175 nom. nud.
*Echinocereus whipplei var. heilii Spelling error for Echinocactus whipplei.
Echinocereus winterianus* Kunzmann 1985:79. nom. nud.
Echinocereus yavapaiensis Baker 2006:63.
  Type - locality: 7 km NNe of Crown King, Yavapai County, Arizona, Baker & Lisenby (ASU).
Echinocereus zuniensis* Eng. & Big. ex Haage & Schmidt 1918 nom. nud. In advertisement opposite p. 74 Monatsschrift für
  Kakteenkunde. 28(7). = Echinocereus mojavensis var. zuniensis.

Abkürzungen & Bibliographie

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Herbarium Abbreviations

(ARIZ) - University of Arizona, Tucson.
(ASU) – Arizona State University.
(B) - Dahlem Botanical Museum, Berlin.
(CA) - California Academy of Sciences, San Francisco.
(DES) - Desert Botanical Garden, Phoenix.
(DS) - Dudley Herbarium, California Academy of Sciences, San Francisco.
(HBG) - University of Hamberg.
(HNT) - Huntington Botanical Garden.
(IBUG) - Universidad de Guadalajara, Zapopan, Jalisco. Mexico.
(JE) - Friedrich - Schiller University, Jena.
(K) - Royal Botanical Gardens, Kew.
(KR) - Staatliches Museum fur Naturkunde Karlsruhe
(LAM) - Natural History Museum of Los Angeles.
(MEXU) - Universidad Autonoma de Mexico.
(MICH) - University of Michigan, Ann Arbor.
(MO) - Missouri Botanical Garden, Saint Louis.
(OKL) - University of Oklahoma, Norman.
(P) - Museum National d’Histoire Naturalle, Paris, France
(POM) - Pomona College, Claremont, California.
(PRC) - Charles University, Prague.
(SRSC) - Sul Ross State College, Alpine, Texas.
(UC) - University of California, Berkeley.
(UNM) - University of New Mexico, Albuquerque.
(US) - United States National Herbarium, Washington, DC.
(WU) - University of Vienna.
(ZSS) - Stadtische Sukkulenten-Sammlung, Zurich.

Other Abbreviations

Auct.? - author unknown.
CSJ - Cactus & Succulent Journal (US).
fa. - forma.
HK - Horst Kunzler Catalog.
Hort. - Hortus (Garden name).
nom. illegit. - name that is illegitimate according to the rules of botanical nomenclature.
nom. inval. - name that is invalid according to the rules of botanical nomenclature.
nom. nud.- a name without any description.
nom. prov. - a provisional name.
pro syn. - as a synonym for
subsp. - subspecies
var. - varietas.
(?) - location of type specimen unknown.

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* indicates work not seen.

1 It is unclear if the subspecific names in Schelle 1907 should be considered forma or varietas. Although he uses the German term
"Formen" he ascribes the names to authors who had published them as varietas. He specifically uses "var." in his 1926 book.

Terry L. Corbett
Las Cruces, NM
18 July 2017