Alle Artikel in der Kategorie “Echinocereus Index

Introduction by Terry L. Corbett

I became interested in the genus Echinocereus in the 1950s when I came upon plants of Echinocereus fendleri and Echinocereus coccineus while hiking in the hills and mountains about Las Cruces, New Mexico. My interest was further stimulated by Dr. Benson’s book, The Cacti of Arizona, Britton and Rose’s Monograph, the Cactus and Succulent Journal and the catalogs of Gates Cactus Garden. The first problem I noticed in my study of the genus was the confusing plethora of names. I began the Echinocereus Index in the early 1960s as an effort to produce a list of all the names of species, subspecies, varieties and forms of the genus Echinocereus. Britton and Rose’s monograph provided a starting point, to which were added names from the Index Kewensis and the Gray Herbarium Card Catalog. Over the intervening years many other sources have been consulted and the index revised several times. Early in 1996 the index was transferred to a computerized file. This file has been circulated to many people through the internet, and late in 1996 the file was added by Martina and Andreas Ohr to their „Echinocereus Homepage.“ Since then, through the response of many Echinocereus enthusiasts, the index has been updated and corrected. Undoubtedly there are still errors and omissions in the index despite considerable effort to eliminate as many of these as possible. In a few cases the original sources have not been available to check the accuracy of the entry. These have been marked in the bibliography with an asterisk. The „Echinocereus Homepage“ file will continue to be updated with future corrections and additions.

Although the genus Echinocereus was not described by Dr. Engelmann until 1848, the first species which belong to the genus were described by Augustin de Candolle in 1828 from collections made by Thomas Coulter in central Mexico. These were Cereus pentalophus and Cereus cinerascens. Other species were described as Echinocactus or Echinopsis. At the suggestion of Prince Salm-Dyck, Engelmann (1849) reduced the genus to the rank of subgenus under Cereus. Labouret (1853) and Coulter (1896) followed Engelmann in placing Echinocereus in the genus Cereus, but Lemaire (1868), Rümpler (1885), Schumann (1897), and Britton and Rose (1922) all keep Echinocereus as a separate genus. Most modern taxonomic treatments also keep Echinocereus as a separate genus. Taylor (1985b) added those plants formerly classed as Wilcoxia to Echinocereus and the index has been revised to reflect this addition. Many people have been helpful in creating this index. Some of those who encouraged my early efforts such as Dr. Edward Castetter, Dudley Gold, Dale Morrical and Dr. Lyman Benson, are no longer with us, but without their help the index project would have been abandoned many years ago. Recently I have had the assistance of many people via the Cactus_ect e-mail list, in particular Lino Di Martino, Andreas and Martina Ohr, Billie Beaston, Bill LaHaye, and Michael Lange. The index has been expanded from just a list of names so that it now includes the author and place of publication for each name and information on the type locality and type specimen. The basionym and a list of synonyms (which are based on the same type specimen, designated by their author as synonyms, or  contain the basionym) have been added so that the index can be used as an organized guide to the many names which have been applied to the plants in this genus. Information relating to other possible synonyms  based on taxonomic interpretations are contained in notes at the end of each entry. The application of taxonomic synonyms is subject to different interpretation by different botanists. The index is intended to be a guide to the different names in this genus for Echinocereus enthusiasts and not a taxonomic treatment of the genus. I have not indicated any preference for any of the different taxonomic systems that have been proposed. Names that do not conform to the standards of botanical nomenclature have been marked with an asterisk. Since the name of a plant includes the name of its author and date it has been necessary in some cases to list the name more than once. In most cases the earlier names are just nom. nud. that probably belong to the later properly published name, but in some cases the different names refer to entirely different plants. I hope this index will be of help to everyone interested in the study, propagation, and preservation of these marvelous plants. The index is alphabetical by species, then subspecific ranks, then date. The order of the data is Genus, species, subspecies/variety/ forma, (basionym author if different) Author, date published:page. = basionym or index base name. TL: [Type locality], collector and type if known, date collected. The abbreviation of the herbarium where the type specimen is deposited and the specimen number are surrounded by ( ). The symbol (?) indicates it is uncertain if a type specimen was preserved. The full citation of the work in which the name was published can be found by checking under the author’s name and date in the bibliography, i.e. Taylor 1985b:120 is page 120 of The Genus Echinocereus. An asterisk* before the genus name means the plant properly belongs to another genus. An asterisk * after other names means the name is invalid or illegitimate. Names in bold are index base names and are followed by information about the basionym and other publication data. For full information of names that are not index base names please check under the name which follows the = sign. Please note that in those cases where the base name is not the basionym the basionym follows the = sign under the bold faced index base name. For the purpose of this index the rank used by the original author has been followed in determining the index base name. In those cases where the original author raised the rank of the plant the higher rank is followed in determining the index base name. This is for reference purposes only and does not indicate any preference in regard to the proper placement of any plant. Names that have never been published under the name Echinocereus are listed with the name of the genus that was used in the basionym. Autonyms (such as Echinocereus acifer var. acifer) are not listed. To find Echinocereus xxx var. xxx fa. yyy look under Echinocereus xxx fa. yyy. Nomenclatural synonyms are listed chronologically.

Las Cruces, New Mexico 1 May 2006. Permission to copy or reprint
for noncommercial purposes is granted.

Terry L. Corbett

Echinocereus abbae – Echinocereus crispatus

Echinocereus Engelmann 1848a:91.

Type: Echinocereus viridiflorus Engelmann
 
Echinocereus abbeae Parsons 1937:6.
TL: Mexico, Sonora, Delta Bacuachic River, 30m (29º49’N,
111º51’W), S. H. Parsons, 2 March 1932 (?).
Synonyms:
Echinocereus fasciculatus ssp. abbeae* Blum 1997 Seed List. nom nud.
       Note:Listed erroneously as a synonym of Echinocereus fenderivar. albiflorus
       (Backeberg 1960:2047, Bravo 1991:70). Taylor (1985b:50) treats this species
as a synonym of Echinocereus fendleri var. fasciculatus.
Echinocereus acifer* Engelmann ex Haage 1859:19 nom. nud.  
[Probably = Echinocereus acifer Jacobi].
Echinocereus acifer (Otto ex Salm-Dyck)Jacobi 1856:109.
= Cereus acifer Otto ex Salm-Dyck 1850:189.
TL: unknown
Synonyms:
Echinocereus longispinus* Hort. ex Schelle 1926:180 pro syn.
        Echinocereus triglochidiatus var. acifer (Otto ex Salm-Dyck)
Bravo 1978:66.
Echinocereus polyacanthus var. acifer* Parker 1996:4
nom. nud.
Echinocereus polyacanthus ssp. acifer (Otto ex Salm-Dyck) Taylor 1997:9.
            Note: Taylor (1984:159) uses the name Echinocereus polyacanthus var. densus
for this species. Cereus acifer* Otto ex Förster 1846:433 nom. nud.is usually
considered a synonym of this species.
Echinocereus acifer* Lemaire 1868:57 nom. nud.
[Probably = Echinocereus acifer Jacobi].
Echinocereus acifer var. brevispinulus Jacobi 1856:109-110.
TL: Mexico.
Echinocereus acifer  var. brevispinus Haage 1859:19 nom. nud.
Echinocereus
acifer var. diversispinus Schumann 1897:287.
TL: Mexico, Coahuila & Durango.
Echinocereus
acifer  var. durangensis (Poselger ex Rümpler)
Schumann 1897:287
= Echinocereus durangensis.
Echinocereus
acifer ssp. huitcholensis (Weber) Lange 1994:109
= Echinocereus huitcholensis.
Echinocereus
acifer  var. longispinus* Haage 1892:116 nom. nud.
Echinocereus
acifer var. tenuispinus Jacobi 1856:109.
TL: North America.
Echinocereus acifer subsp. topiensis (W.Rischer & Trocha) W.Rischer Echinocereenfreund 17(4): 88. 2004.
   = Echinocereus topiensis W.Rischer & Trocha  1999

Echinocereus acifer var. trichacanthus Hort. ex Hildmann 1891:44.
TL: unknown
Echinocereus acifer var. tubiflorus Rischer in Blum et al. 1998:409-410.
  TL: Mexico, Zacactecas – Durango border, San Juan Capistran, Rischer
017,
April 1990 (ZSS AX 17983).
Cereus acinaciformis* Steudel 1841:333 pro syn.
= Echinocereus cinerascens.
Cereus
acinaciformis* Hort. Berol. ex Salm-Dyck 1845:27 pro syn.
= Echinocereus cinerascens.
Cereus
aciniformis* Otto 1833:336 nom. nud.
= Echinocereus cinerascens
var. crassior [According to
Pfeiffer 1837:101].
Cereus aciniformis* Hort. Berol. ex Pfeiffer 1837:101 pro syn.
= Echinocereus cinerascens
var. crassior.
Echinocereus adustus Engelmann 1848a:104.
TL: Chihuahua, Cosihuiriachi, 1900m, Wislizenus, 1846. (MO).
Synonyms:
Cereus adustus (Engelmann) Engelmann 1849:50.
Echinocereus pectinatus var. adustus (Engelmann) Daul 1890:78.
Echinocereus pectinatus fa. adustus (Engelmann) Voss 1894:378.
Echinocereus pectinatus var. adustus* Schumann 1897:271.
Echinocereus caespitosus var. adustus (Engelmann)
Engelmann ex Borg 1937:174.
   Note: Echinocereus radians and Echinocereus rufispinus are usually
considered synonyms of this species.
Cereus adustus var. radians (Engelmann) Engelmann 1849:50 = Echinocereus radians.
Echinocereus adustus  subsp. roemerianus W.Rischer, Echinocereenfreund 16(2): 44 (42-50; figs.). 2003
TL: Canatlan, MX, R. C. Roemer 6066714 (UNAM)
Echinocereus adustus var. schwarzii (Lau) Taylor 1985a:268 = Echinocereus schwarzii.
Echinocereus
adustus ssp. schwarzii (Lau) Taylor 1997:8  = Echinocereus schwarzii.
Echinocereus adustus var. schwarzii fa. madrensis* Parker 1996:6.nom. nud.
= Echinocereus madrensis.
Echinocereus
adustus var. schwarzii fa. radians* Parker 1996:6.  nom. nud.
= Echinocereus radians.
Echinocereus
adustus var. schwarzii fa. rufispinus* Parker 1996:6. nom. nud.
= Echinocereus rufispinus.
Echinocereus aggregatus
* Rydberg 1906:146 nom. illegit.
= Mammillaria aggregata*
Engelmann 1848b:157. nom. prov.
  = Mammillaria aggregata Engelmann ex  Watson 1878:398.
  [Rydberg used this name instead of Echinocereus coccineus].
see Echinocereus coccineus ssp. aggregatus.
Echinocereus
aguirrei* Glass 1965:160 nom. nud.  [a form of Echinocereus pulchellus].
Echinocereus albatus* Backeberg 1960:2007-2008 nom. inval. (no type designated).
    Synonyms:
Echinocereus albispinus* Hort. ex Backeberg 1960:2007 pro syn.
Echinocereus longisetus
var. albatus* Sterk 1980:12-13 nom. inval.
       Note:Usually considered a synonym of Echinocereus nivosus.
Echinocereus albiflorus Weingart 1933:156.
TL: Mexico, Chihuahua, Halbinger (?).
    Synonyms:
Echinocereus fendleri var. albiflorus (Weingart)
Backeberg 1960:2047.
Echinocereus fendleri fa. albiflorus* Parker 1996: 2 nom. nud.
         Note: Backeberg 1960:2047 and Bravo 1991:70 erroneously
list Echinocereus abbeae as a synonym of this species.
Taylor (1985b) lists this species as a synonym of Echinocereus fendleri.
Echinocereus
albiflorus* Lowry 1937:20 nom. nud.
  [Probably a white flowering form of Echinocereus enneacanthus].
Echinocereus albispinus Lahman 1935:143.
TL: Oklahoma, Medicine Park, Lahman (MO).
Synonyms:
Echinocereus baileyi var. albispinus (Lahman) Backeberg 1941:4.
Echinocereus longispinus var. albispinus* Backeberg 1960:2010.
Echinocereus reichenbachii var. albispinus (Lahman) Benson 1969b:127.
Echinocereus reichenbachii var. baileyi fa. albispinus* Mesa Garden Seed
Catalog 1996 nom. nud.
          Note: The name Echinocereus reichenbachii var. albispinus is used for
          Echinocereus baileyi by Benson (1969b:127). Taylor (1985b) lists this
species as a synonym of Echinocereus reichenbachii var. baileyi.
Echinocereus
albispinus* Hort. ex Backeberg 1960:2007 pro syn.
= Echinocereus albatus.
Echinocereus
amoenus (Dietrich) Weber ex Haage 1892:116.
= Echinopsis amoena Dietrich 1844:187-188.
TL: Mexico, Hidalgo, near San Mateo 7500 ft., Ehrenberg (?).
Synonyms:
Echinopsis pulchella var.flore kermesina* Haage ex Förster1846:364. pro syn.
        Echinopsis pulchella var. amoena (Dietrich) Förster 1846:364.
        Echinopsis pulchella var. rosea* Labouret 1853:292. pro syn.
        Cereus amoenus (Dietrich) Hemsley 1880:540.
Echinocereus pulchellus var. amoenus (Dietrich)Förster ex
Schumann 1897:253.
Echinocereus pulchellus ssp. amoenus (Dietrich)Blum 1997 Seed
list. nom. nud.
            Note: Echinocereus amoenus is usually attributed to Schumann 1894:185.
Taylor (1989:75) notes that the plant usually known in cultivation as
Echinocereus amoenus“ is instead a form of Echinocereus weinbergii.
Echinocereus
amoenus var. aguirre* Zehnter & Künzler Catalog nom. nud. 
= Echinocereus aguirrei*.
Echinocereus angusticeps Clover 1935:79.
TL: Texas, Hidalgo Co., Linn , Clover 15261 (MICH).
Synonyms:
Echinocereus papillosus var. angusticeps (Clover) Marshall & Bock 1941:119.
Echinocereus blanckii var. angusticeps (Clover) Benson 1969b:260.
Echinocereus berlandieri var. angusticeps (Clover)Benson 1976:59.
Echinocereus papillosus ssp. angusticeps* Blum 1997 Seed List nom. nud.
          Note: Benson uses the names Echinocereus blanckii var. angusticeps and
          Echinocereus berlandieri var. angusticeps for Echinocereus papillosus.
Echinocereus
anneacanthus* Daul 1890:73.
TL: unknown
Echinocereus apachensis Blum & Rutow in Blum et al. 1998:101.
TL: Arizona,Maricopa Co. Fish Creek, Mc Kelvey 735, 18 Feb. 1929
(US 1532949).
Echinocereus arizonicus Rose ex Orcutt 1926:3.
TL: Arizona, Boundary Monument between Pinal & Gila Counties,
4700 ft., C. R. Orcutt, July 1922 (NY).
    Synonyms:
Echinocereus triglochidiatus var. arizonicus (Rose ex Orcutt)
Benson 1969a:21.
Echinocereus coccineus var. arizonicus (Rose ex Orcutt)
Ferguson 1989:221.
Echinocereus arizonicus ssp. matudae (Bravo) Rutow in Lange & Rutow 1994:51
=
Echinocereus matudae.
Echinocereus arizonicus
ssp. nigrihorridispinus Blum & Rutow in Blum et al.
1998:370-371.
TL: New Mexico, Luna Co., HK 1054 (ZSS AX 17982).
Echinocereus armatus (Poselger) Poselger ex Backeberg &
Knuth 1935:318
    = Echinocereus pectinatus var. armatus.
*Echinocereus arrojadei Löfgren
[from Brazil and therefor not an Echinocereus].
Echinocereus australis* Hort. ex Backeberg 1960:2045 pro syn.
= Echinocereus fendleri.
Echinocereus
baileyi Rose 1909b:403.
TL: Oklahoma, Wichita Mtns., V. Bailey, Aug. 1906 (US 53167).
Synonyms:
Echinocereus reichenbachii var. baileyi (Rose) Taylor 1985b:133.
      Echinocereus reichenbachii ssp. baileyi (Rose) Taylor 1997:9.
Note: The name Echinocereus reichenbachii var. albispinus is used for
this species by Benson (1969b:127 ).
Echinocereus baileyi var. albispinus (Lahman) Backeberg 1941:4
    = Echinocereus albispinus Lahman.
Echinocereus baileyi var. brunispinus Backeberg 1941:4.
TL: Oklahoma, Wichita Mtns.
Note: Backeberg 1960:2010 lists Echinocereus longispinus and Echinocereus
oklahomensis
as synonyms of this variety.
Echinocereus baileyi var. caespiticus* Backeberg 1960:2011 nom. inval.
(no type designated).
Echinocereus
baileyi var. flavidispinus Backeberg 1941:4.
TL: Oklahoma, Wichita Mtns.
Echinocereus baileyi var. roseispinus Backeberg 1941:4.
TL: Oklahoma, Wichita Mtns.
Echinocereus barcelona* Orcutt 1926:4 nom. nud.
Echinocereus
barcena* Rebut ex Hirscht 1896:126 nom. nud.
Echinocereus barcena Fobe 1911:56.
TL: unknown [from central Mexico according to Borg 1937:178].
Synonyms:
Echinocereus parkeri fa. barcena* Parker 1996:3 nom. nud.
Echinocereus barcena* Rebut ex Berger 1929 nom. illegit.  (later homonym) [Perhaps
= Echinocereus barcena
Fobe].
Echinocereus barkeanus* Heyer 1989:25 nom. nud. [ = Lau 1101].
= Echinocereus rayonesensis.
Echinocereus
barthelowanus Britton & Rose 1922:41.
TL: Baja California,Isla Magdalena, Santa Maria Bay, J. N. Rose,
  18 March 1911 (US 16278).
Echinocereus bergeanus* Haage 1900:156 nom. nud.
Echinocereus
berlandianus* Orcutt 1926:3  error in spelling for Echinocereus berlandieri.
Echinocereus berlandieri (Engelmann) Haage 1859:19.
= Cereus berlandieri Engelmann 1856:286.   [fuller description Engelmann 1859:38, t.58].
TL: Texas, Nueces River, Berlandier 2423, April 1834 (MO).
Synonyms:
Echinocereus blankii* Hort. ex Palmer 1865:92 pro syn.
        Echinocereus blanckii var. berlandieri (Engelmann) Backeberg 1960:1999.
Note: Echinocereus poselgerianus is usually considered a synonym of this species.
This species is often erroneously treated as a synonym of Echinocereus blanckii.
Echinocereus
berlandieri var. angusticeps (Clover) Benson 1976:59
     = Echinocereus angusticeps.
Echinocereus
berlandieri var. blanckii* Fournier 1935:21 nom. illegit.
= Echinocereus blanckii
.
Echinocereus berlandieri var. longispinus* Hort. ex Schelle 1926: 163 nom. nud.
Echinocereus
berlandieri var. papillosus (Linke ex Rümpler) Benson 1976:59
     = Echinocereus papillosus.
Echinocereus
berlandieri var. poselgerianus (Linke) Lode 1992:17
= Echinocereus poselgerianus.
*Echinocereus bertinii = Austrocactus bertinii.
*Echinocereus bicolor  Galeotti = error in spelling for Echinocactus bicolor.
Cereus
bigelovii* Engelmann 1857: t. 4 nom. nud. = Echinocereus mojavensis.
Cereus bigelovii
var. zuniensis* Engelmann 1857: t. 4 nom. nud.
= Echinocereus mojavensis
var. zuniensis.
Echinocereus
blanckii (Poselger) Poselger ex Rümpler 1885:779.
= Cereus blanckii Poselger 1853:134.
TL:Mexico, Tamaulipas, near Camargo, Poselger (?).
Synonyms:
Echinocereus berlandieri var. blanckii* Fournier 1935:21 nom. illegit.
            Note: Taylor (1985b) lists this species as a synonym of Echinocereus
enneacanthus
var. brevispinus. This name has been errroneously applied
to Echinocereus berlandieri.
Echinocereus
blanckii var. angusticeps (Clover) Benson 1969b:260
     = Echinocereus angusticeps.
    Note: This is the name Benson uses for Echinocereus papillosus.
Echinocereus
blanckii var. berlandieri (Engelmann) Backeberg 1960:1999
     = Echinocereus berlandieri.
Echinocereus
blanckii var.inermis* Heyer 1989:26 pro syn.
     = Echinocereus moricallii.
Echinocereus
blanckii var. leonensis (Mathsson) Backeberg 1960:1999
     = Echinocereus leonensis.
Echinocereus
blanckii var. papillosus (Linke ex Rümpler) Benson 1969d:126
     = Echinocereus papillosus.
Echinocereus
blanckii var. poselgerianus (Linke) Backeberg 1960:1999
     = Echinocereus poselgerianus.
Echinocereus
blankii* Hort. ex Palmer 1865:92 pro syn.
= Echinocereus berlandieri.
Echinocereus
bolansis* Runge ex Haage 1892:117 pro syn.
= Echinocereus conglomeratus.
*Echinocereus boliviensis Poselger ex Schumann 1897:290.
  [from Bolivia and therefor not an Echinocereus].
Echinocereus bonatzii Römer 1995:98-113.
TL: Durango, Sierra Mezquital, H. J. Bonatz 12388 (ZSS).
Synonyms:
Echinocereus pamanesiorum ssp. bonatzii (Römer) Römer 1997:45.
Echinocereus bonkerae Thornber & Bonker 1932:71-73, 85.
TL: Arizona, Pinal Mtns. near Oracle, Thornber & Bonker   (ARIZ 156240).
Synonyms:
Echinocereus fendleri var. bonkerae (Thornber & Bonker) Benson 1944:260.
      Echinocereus boyce-thompsonii var. bonkerae (Thornber & Bonker)  Peebles 1949:191-192 .
      Echinocereus fasciculatus var. bonkerae (Thornber & Bonker)  Benson 1969a:21.
Echinocereus engelmannii var. bonkerae* Mesa Garden Seed Catalog 1996. nom. nud.
      Echinocereus fasciculatus ssp. bonkerae  (Thornber & Bonker) Taylor 1997:8.
Echinocereus boyce-thompsonii Orcutt 1926:4.
TL: Arizona, Boyce-Thompson Arboretum, Orcutt (?).
Synonyms:
Echinocereus fendleri var. boyce-thompsonii (Orcutt) Benson 1944:260.
Echinocereus fasciculatus var. boyce-thompsonii (Orcutt) Benson 1969a:22.
Echinocereus engelmannii var. boyce-thompsonii* Mesa Garden Seed Catalog 1996 nom. nud.
      Echinocereus fasciculatus ssp. boyce-thompsonii (Orcutt) Taylor 1997:8.
Echinocereus boyce-thompsonii var. bonkerae (Thornber & Bonker) Peebles 1949:191-192
     = Echinocereus bonkerae.
Echinocereus
brandegeei (Coulter) Coulter ex Schumann 1897:290.
= Cereus brandegeei Coulter 1896:389.
TL: Baja California, between El Campo Aleman & San Gregorio, T.S. Brandegee, 1889 (UC).
Note: Echinocereus mamillatus and Echinocereus sanborgianus are usually
considered synonyms of this species.
Echinocereus brandegeei  fa. mamillatus* Parker 1996:1 nom. nud.
= Echinocereus mamillatus.
Echinocereus
brandegeei  var. sanborgianus* Lode 1994: no. 1272 nom. inval.
= Echinocereus sanborgianus.
Echinocereus
brandegeei  fa. sanborgianus* Parker 1996:1 nom. nud.
     = Echinocereus sanborgianus.
Echinocereus
brenbergii* Lowry 1933:410 nom. nud.
Echinocereus
bristolii Marshall 1938:160-161.
TL: Sonora, Soyopa, 5000 ft., Bristol & Marshall, 1934 (DS 251175).
Synonyms:
Echinocereus pectinatus var. bristolii (Marshall) Marshall 1956b:81.
Echinocereus bristolii ssp. floresii (Backeberg) Blum & Lange in Blum et al. 1998:353. 
= Echinocereus floresii.
Echinocereus
bristolii var. pseudopectinatus Taylor 1985b:120
[see Echinocereus pseudopectinatus].
Echinocereus
caespitosus (Engelmann) Engelmann 1848a:110.
= Cereus caespitosus Engelmann 1845:247.
TL: Texas, Austin Co., Near Cat Springs, Lindheimer,
    June 1845 (MO).
Synonyms:
Cereus caespitosus var. minor* Engelmann 1856:280[Based on
same type as Echinocereus caespitosus].
Echinocereus pectinatus var. caespitosus (Engelmann) Schumann 1897:272.
Echinocereus caespitosus var. minor* Engelmann ex Weniger 1970:22 nom. inval.
        Echinocereus reichenbachii fa. caespitosus* Parker 1996:5 nom. nud.
        Echinocereus reichenbachii var. caespitosus* Mesa Garden Seed Catalog 1996 nom. nud.
        Echinocereus reichenbachii ssp. caespitosus (Engelmann) Blum & Lange in Blum et al. 1998:324.
Note: Echinocereus caespitosus is usually treated as a synonym of Echinocereus reichenbachii.
Echinocereus
caespitosus var. adustus (Engelmann) Engelmann ex Borg 1937:174.
     = Echinocereus adustus.
Echinocereus
caespitosus var. armatus (Poselger) Poselger ex Borg 1937:174
     = Echinocereus pectinatus var. armatus.
Echinocereus
caespitosus var. candicans* Hort. ex Borg 1937:174 pro syn.
= Echinocereus reichenbachii.
Echinocereus
caespitosus var. castaneus (Engelmann) Engelmann ex Rümpler 1885:811.
= Cereus caespitosus var. castaneus Engelmann 1850:203.
TL: Texas, Granitic region of the Llano, Lindheimer, 1847 (MO).
        Synonyms:
Cereus reichenbachii var. castaneus (Engelmann) Labouret 1853:319.
Echinocereus pectinatus var. castaneus (Engelmann) Daul 1890:78.
Echinocereus castaneus (Engelmann) Orcutt 1926:3.
Echinocereus pectinatus var. jamauvei *Fric ex Orcutt 1926:4 pro syn.
          Echinocereus pectinatus var. adustus fa. castaneus (Engelmann) Engelmann ex Schelle 1926:174.
Echinocereus pectinatus fa. castaneus (Engelmann) Krainz 1967:50.
Echinocereus caespitosus var. chrysacanthus (Schumann) Schumann ex Borg 1937:174
    = Echinocereus pectinatus var. chrysacanthus.
Echinocereus
caespitosus var. major (Engelmann) Engelmann ex Rümpler 1885:811.
= Cereus caespitosus var. major Engelmann 1856:280.
TL: Texas, New Braunfels, Lindheimer, 1851(MO).
Echinocereus caespitosus var. minor* Engelmann ex Weniger 1970:22 nom. inval.
    = Cereus caespitosus var. minor* Engelmann 1856:280new name for Echinocereus
caespitosus
var. caespitosus.
Echinocereus caespitosus var. perbellus* Weniger 1970:23 nom. inval. (no basionym given)
= Echinocereus perbellus.
Echinocereus
caespitosus var. purpureus* Weniger 1970:23 nom. inval. (no basionym given)
= Echinocereus purpureus.
Echinocereus
caespitosus var. reichenbachii* Ters. ex Borg 1937:174 nom. illegit.
= Echinocereus reichenbachii.
Echinocereus
caespitosus var. rubescens* Hort. ex Borg 1951:221 nom. inval (no Latin diagnosis).
Echinocereus
caespitosus var. rufispinus (Schumann) Schumann ex Borg 1951:221
= Echinocereus pectinatus var. rufispinus.
Echinocereus
caespitosus var. tamaulipensis (Fric) Fric ex  Borg 1937:174 
= Echinocereus pectinatus
var. tamaulipensis.
*Echinocereus candicans Rümpler 1885 = Echinopsis candicans.
*Echinocereus candicans var. tenuispinus = Echinopsis candicans
Echinocereus
candicans* Rünge ex Haage 1859:19 pro syn.
= Echinocereus pectinatus
var. rigidissimus.
Echinocereus
canyonensis Clover & Jotter 1941:417-418.
TL: Arizona, Coconino Co., Bass Cable, Grande Canyon, Clover & Jotter 2317 (MICH).
Synonyms:
Echinocereus coccineus ssp. canyonensis* Blum 1997 Seed List nom. nud.
          Note: Taylor (1985b) lists this species as a synonym of Echinocereus triglochidiatus
var. melanacanthus.
Echinocereus carmenensis
Blum, Lange & Scherer in Blum et al. 1998:221.
TL:Mexico, Coahuila, Sierra del Carmen, SB 1476 (ZSS A 17986).
Echinocereus carnosus Rümpler 1885:796-797.
TL: Texas
Synonyms:
Echinocereus enneacanthus var. carnosus (Rümpler) Schelle 1907:127.
Echinocereus enneacanthus var. brevispinus fa. carnosus* Parker 1996:2 nom. nud.
Echinocereus castaneus*
Engelmann ex Haage 1859:19nom. nud.
Echinocereus
castaneus (Engelmann) Orcutt 1926:3 = Echinocereus caespitosus  var. castaneus.
*Echinocereus centralis Rose = Echinomastus centralis.
Echinocereus
cereiformis* Auct. ? (Kakteenkunde 1935:1) nom. inval. (no Latin diagnosis).
Note: Taylor (1985b) lists this name as a synonym of Echinocereus berlandieri.
Echinocereus
cernosus* Fric 1929:24 nom. nud.  [Probably = Echinocereus pectinatus].
Echinocereus
chihuahuensis* Shields 1961:4. nom. inval. (no Latin diagnosis)
*Echinocereus chiloensis = Echinopsis chiloensis.
Echinocereus
chisoensis Marshall 1940:15.
TL: Texas, Chisos Mtns., F. Radley, 10 Apr. 1939 (DS 263216).
Synonyms:
Echinocereus reichenbachii var. chisoensis (Marshall) Benson 1969d:127.
Echinocereus
chisoensis var. fobeanus (Oehme) Taylor 1985a:261= Echinocereus fobeanus.
Echinocereus
chisoensis var. metornii* Parker 1996:5 nom. nud. = Echinocereus metornii.
Echinocereus
chloranthus (Engelmann) Haage 1859:19.
= Cereus chloranthus Engelmann 1856:278-279. [fuller description Engelmann 1859:29].
TL: Texas, stony hills near Frontera [El Paso], Wright 95, 2 Apr.
1852 (MO 2016809).
      Synonyms:
Echinocereus viridiflorus var. chloranthus (Engelmann) Backeberg 1960:2015.
Echinocereus viridiflorus fa. chloranthus (Engelmann) Krainz 1967:51.
Echinocereus viridiflorus ssp. chloranthus (Engelmann) Taylor 1997:10.
Note: Usually listed as Echinocereus chloranthus Engelmann ex Rümpler 1885:814.
Echinocereus chloranthus var. albispinus* Fric 1924:121 nom. nud.
Echinocereus
chloranthus var. cylindricus (Engelmann) Taylor 1984:169
      = Echinocereus viridiflorus var. cylindricus.
Echinocereus
chloranthus ssp. cylindricus Blum & Lange in Blum et al. 1998: 205.
= Echinocereus viridiflorus var. cylindricus.
Echinocereus
chloranthus var. flaviflorus* Hort. nom. nud.
Echinocereus
chloranthus var. flavispininus* Ito 1952:133 nom. inval (no Latin diagnosis).
Echinocereus
chloranthus var. longisetus* Hort. nom. nud.
Echinocereus
chloranthus var. livermorensis*  Kolle 1978:34 nom inval.
= Echinocereus viridiflorus
var. weedinii.
Echinocereus chloranthus var. milleri* Höchstätter Seed List 1997 nom. nud.
[See Echinocereus milleri*].
Echinocereus chloranthus var. neocapillus Weniger 1969:39-41.
TL: Texas, 5 to 10 miles south of Marathon, Weniger 711 (UNM).
    Synonyms:
Echinocereus viridiflorus var. neocapillus (Weniger) Leuck 1980:35-36.
Echinocereus russanthus var. neocapillus* Mesa Garden Seed Catalog 1996 nom. nud.
      Echinocereus neocapillus Blum & Lange in Blum et al. 1998:219.
Echinocereus chloranthus var. rhyolithensis  Blum & Lange in Blum et al. 1998:208.
TL: New Mexico, Sierra County, Lake Valley,  Pierce 110,  1961 (UNM).
Echinocereus chloranthus var. roseiflorus* Ito 1981.(according to Evanich 1989:103) nom. nud.
Echinocereus
chloranthus var. russanthus (Weniger) Lamb ex Rowley 1974a:7
      = Echinocereus russanthus.
Echinocereus chloranthus var. senilis* Fric 1924:121 nom. nud.
Echinocereus
chloranthus var. vulpis-cauda* Chudleigh 1967:43 nom. nud. 
[Probably = Echinocereus russanthus].
Echinocereus
chloranthus ssp. weedinii* Höchstätter Seed List 1997 nom. nud. 
[See Echinocereus viridiflorus var. weedinii].
Echinocereus chlorophthalmus (Hooker) Britton & Rose 1913:242.
= Echinocactus chlorophthalmus Hooker 1848:t. 4373.
TL: Mexico, Hidalgo, Real del Monte.
Synonyms:
Echinocereus cinerascens fa. chlorophthalmus* Parker 1996:2 nom. nud.
            Note: This species is often considered a synonym of Echinocereus cinerascens.
Echinocereus
chrysocentrus (Engelmann & Bigelow) Orcutt 1926:4
= Echinocereus engelmannii
var. chrysocentrus.
Echinocereus
cinerascens* Engelmann  ex  Haage 1859:19 nom. nud. 
[Probably = Echinocereus cinerascens].
Echinocereus cinerascens* Lemaire 1868:56 nom. nud. 
[Probably = Echinocereus cinerascens].
Echinocereus cinerascens (De Candolle) De Candolle ex
Rümpler 1885:786.
= Cereus cinerascens De Candolle 1828:116.
TL: Mexico, Zimapan, Coulter 23 (?).
Synonyms:
          Cereus acinaciformis* Steudel 1841:333 pro syn.
          Cereus acinaciformis* Hort. Berol. ex Salm-Dyck 1845:27 pro syn.
          Echinocereus deppei*(Salm-Dyck) Schumann 1895:123 pro syn.
Echinocereus
undulatus*Hildmann ex Schumann 1895:124 pro syn.
              Note: Cereus deppei, Echinocereus cirrhiferus and Echinocereus glycimorphus 
are usually considered synonyms of Echinocereus cinerascens.
Echinocereus
cinerascens var. caesius* Hort. ex Schelle 1926:169 nom. nud.
Echinocereus
cinerascens fa. chlorophthalmus* Parker 1996:2 nom. nud.
= Echinocereus chlorophtahlmus
Echinocereus
cinerascens var. crassior (De Candolle) De Candolle ex Rümpler 1885:786-787.
= Cereus cinerascens var. crassior De Candolle 1828:116.
TL: Mexico, Zimapan, Coulter (?).
Synonyms:
Cereus aciniformis* Otto 1833:336 nom. nud.
          Cereus aciniformis* Hort. Berol. ex Pfeiffer 1837:101 pro syn.
Echinocereus
cinerascens var. ehrenbergii (Pfeiffer) Bravo 1974:47
      = Echinocereus ehrenbergii.
Echinocereus
cinerascens ssp. ehrenbergii (Pfeiffer)Blum & Rutow in Blum et al. 1998:140-141.
= Echinocereus ehrenbergii.
Cereus
cinerascens var. fendleri (Engelmann) Bois 1928:371
     = Echinocereus fendleri.
Echinocereus
cinerascens fa. glycimorphus* Parker 1996:2 nom. nud.
= Echinocereus glycimorphus.
Echinocereus
cinerascens var. schmidtii* Haage 1859:19 nom. nud.
Echinocereus
cinerascens var. septentrionalis Taylor 1988:68-69.
TL: Mexico, San Luis Potosi, Entronque Huizache, Taylor 281, 10
July 1986 (MEXU).
Echinocereus
cinerascens  ssp. septentrionalis Taylor 1997:8
= Echinocereus cinerascens var. septentrionalis .
Echinocereus cinerascens fa. spinibarbis* Parker 1996:2 nom. nud.
= Echinocereus spinibarbis*
Hildmann[not Salm-Dyck].
Echinocereus cinerascens var. tenuior (De Candolle) De Candolle ex Rümpler 1885:776.
= Cereus cinerascens var. tenuior De Candolle 1828:116.
TL: Mexico, Zimapan, Coulter (?).
Echinocereus cinerascens var. tulensis Taylor 1988:69 = Echinocereus tulensis.
Echinocereus cinerascens ssp. tulensis  (Bravo) Taylor 1997:8 = Echinocereus tulensis.
     = Echinocereus tulensis.
*Echinocereus cinnabarinus = Echinopsis cinnabarina.
Echinocereus cirrhiferus
* Engelmann ex Haage 1859:19 nom. nud.
Echinocereus
cirrhiferus (Labouret) Labouret ex Rümpler 1885:778.
= Cereus cirrhiferus Labouret 1853:311.
TL: Mexico.
Note: Usually considered a synonym of Echinocereus cinerascens.
*Echinocereus clavatus  = Eridisia spiniflora.
Echinocereus claviformis*
Haage 1859:19. nom. nud.
Echinocereus
claviformis (Regel & Klein) Haage 1893:22
= Cereus claviformis Regel & Klein 1860:46.
TL: Mexico, Karwinsky ( ? ).
Echinocereus coccineus Engelmann 1848a:93.
TL: New Mexico, Wolf Creek, Wislizenus,24 June 1846 (MO).
    Synonyms:
Cereus coccineus (Engelmann) Engelmann 1849:51 nom. illegit. 
[not Cereus coccineus Salm- Dyck 1828].
Echinocereus phoeniceus* Engelmann ex Rümpler 1885:788 nom. illegit.
= Cereus phoeniceus
Engelmann 1856:284.
Echinocereus melanacanthus* Engelmann ex Earl 1963:61 printing error.
      Echinocereus triglochidiatus var. coccineus (Engelmann) Engelmann ex Marshall & Bock 1941:117.
Note: Benson uses the name Echinocereus triglochidiatus var. melanacanthus for this species.
Echinocereus coccineus ssp. aggregatus
Blum, Lange & Rutow in Blum et al. 1998:428.
    = Mammillaria aggregata* Engelmann ex Watson 1878:398. pro syn.
   TL: Headwaters of the Gila, 6,000 ft. Emory,18 Oct. 1846 (?)
= Mammillaria aggregata*
Engelmann 1848b:157. nom. prov.
        [See Echinocereus aggregatus].
Echinocereus
coccineus var. arizonicus (Orcutt) Ferguson 1989:221
     = Echinocereus arizonicus.
Echinocereus
coccineus var. arizonicus fa. matudae* Parker 1996:4 nom. nud.
= Echinocereus matudae.
Echinocereus
coccineus ssp. canyonensis* Blum 1997 Seed List nom. nud.
= Echinocereus canyonensis.
Echinocereus
coccineus var. conoideus* Weniger 1970:42 nom. inval.  (no basionym given)
= Echinocereus conoideus.
Cereus
coccineus var. cylindricus Engelmann 1849:51.
TL: New Mexico, Mtns. near Santa Fe, Fendler, 17 Sept. 1846 (MO).
Note: This plant is usually considered as a synonym of Echinocereus coccineus var.
    coccineus and has never been recombined with the name Echinocereus.
Echinocereus coccineus var. gurneyi (Benson) Heil & Brack 1988:26
     = Echinocereus triglochidiatus var. gurneyi.
Echinocereus
coccineus ssp. gurneyi* Blum 1997 Seed List nom. nud.
= Echinocereus triglochidiatus
var. gurneyi .
Echinocereus coccineus fa. hexaedrus* Parker 1996:4 nom. nud.
= Echinocereus hexaedrus.
Echinocereus coccineus
var. inermis (Schumann) Purpus 1925:49.
= Echinocereus phoeniceus var. inermis Schumann 1896a:150.
TL: Colorado, Mesa Grande, Tongue Creek, C. A. Purpus (?).
Synonyms:
Echinocereus triglochidiatus var. inermis (Schumann) Arp  1973:132-133.
Echinocereus triglochidiatus var. mojavensis fa. inermis  (Schumann) Ferguson 1989:219.
Echinocereus triglochidiatus var. melanacanthus fa. inermis* Alabaster 1994:105 nom. inval.
        Echinocereus coccineus var. mojavensis fa. inermis* Parker 1996:4 nom. nud.
        Echinocereus mojavensis fa. inermis* Mesa Garden Seed Catalog 1996 nom. nud.
        Echinocereus triglochidiatus ssp. mojavensis fa. inermis Blum in Blum et al. 1998:365.
Echinocereus coccineus var. kunzei (Gürke) Backeberg 1960:2070
     = Echinocereus kunzei.
Cereus
coccineus var. melanacanthus Engelmann 1849:51.
TL: New Mexico, Santa Fe, Fendler, 26 Nov. 1846 (MO).
    Synonyms:
Echinocereus triglochidiatus var. melanacanthus (Engelmann)
Benson 1944:254.
Note:This is the name Benson applies to Echinocereus coccineus.
Echinocereus coccineus fa. lloydii* Parker 1996:4 nom. nud.
= Echinocereus lloydii.
Echinocereus coccineus
var. mojavensis* Parker 1996:4 nom. nud.
     = Echinocereus mojavensis.
Echinocereus coccineus
ssp. mojavensis (Engelmann & Bigelow) Taylor 1997:8
     = Echinocereus mojavensis.
Echinocereus
coccineus var. mojavensis fa. inermis* Parker 1996:4 nom. nud.
= Echinocereus
coccineus var. inermis.
Echinocereus
coccineus var. neomexicanus* Parker 1996:4 nom. nud.
= Echinocereus neomexicanus.
Echinocereus
coccineus var. neomexicanus fa. rosei* Parker  1996:4 nom. nud.
= Echinocereus rosei.
Echinocereus
coccineus var. nudum* Auct ? (CSJ 35:194) nom. nud.
Echinocereus
coccineus var. octacanthus* Boissevain & Davidson 1940:39 nom. illegit.
= Echinocereus octacantus
.
Echinocereus coccineus var. paucispinus (Engelmann) Ferguson 1989:222
     = Echinocereus paucispinus.
Echinocereus coccineus ssp. paucispinus Blum, Lange & Rutow inBlum et al. 1998:433.
    = Echinocereus paucispinus.
Echinocereus
coccineus fa. roemeri* Parker 1996:4 nom. nud.
     = Echinocereus roemeri.
Echinocereus
coccineus ssp. roemeri Blum, Lange & Rutow in Blum et al. 1998:426.
     = Echinocereus roemeri.
Echinocereus
coccineus fa. rosei* Blum 1990:13. nom. inval. (no basionym given).
= Echinocereus rosei.
Echinocereus
coccineus ssp. rosei Blum & Rutow in Blum et al. 1998:426.
     = Echinocereus rosei.
Cereus
concolor Schott1857:ii .
TL: Texas, Pecos County, Escondido Springs
[Apparently some form of Echinocereus viridiflorus or  Echinocereus chloranthus that
has never been formally transfered to the genus Echinocereus].
Echinocereus congestispinus*  Engelmann exHaage 1859:19 nom. nud.
Echinocereus
congestispinus*  Förster exSeitz 1870:10 nom. nud.
Echinocereus
conglomeratus* Förster exSeitz 1870:10 nom. nud.
Echinocereus
conglomeratus Blanc 1890:56.
TL: Nuevo Leon, Rinconada, Mathsson, 24 Sept. 1889 (?)[according
to Schumann 1897:278].
Synonyms:
Echinocereus bolansis* Runge ex Haage 1892:117 pro syn.
       Cereus conglomeratus (Blanc) Berger 1905:81.
Echinocereus enneacanthus var. conglomeratus (Blanc) Benson 1974:80.
Echinocereus stramineus var. conglomeratus (Blanc) Bravo1974:47.
Echinocereus stramineus fa. conglomeratus* Parker 1996:3 nom. nud.
           Note: This is usually listed as Echinocereus conglomeratus Förster ex Schumann 1897:278.
Echinocereus conglomeratus var. robustior* Haage 1892:117 nom. nud.
Echinocereus
conoideus (Engelmann & Bigelow) Engelmann ex
Rümpler 1885:807.
= Cereus conoideus Engelmann & Bigelow in Engelmann 1856:284.
[fuller description Engelmann 1857:35].
TL: New Mexico, Anton Chico, J. M. Bigelow, 24 Sept. 1853 (MO).
Synonyms:
Cereus phoeniceus ssp. conoideus (Engelmann & Bigelow) Engelmann 1857:37.
Echinocereus phoeniceus* var. conoideus Schumann 1896:150.
Echinocereus coccineus var. conoideus* Weniger 1970:42 nom. inval.
              Note: This species is usually treated as a synonym of  Echinocereus coccineus.
Echinocereus cowperi*
Hort ex  Blum et al. 1998:208 pro syn. Echinocereus
chloranthus ssp.
rhyolithensis.
Echinocereus crispatus* Mathsson 1890:464 nom. nud.

Las Cruces, New Mexico 1 May 2006. Permission to copy or reprint
for noncommercial purposes is granted.

Echinocereus ctenoides – Echinocereus monacanthus

Echinocereus ctenoides* Lemaire 1868:57 nom. nud.
  [Probably = Echinocereus ctenoides Engelmann ex Rümpler].
Echinocereus ctenoides (Engelmann) Engelmann ex Rümpler 1885:819.
= Cereus ctenoides Engelmann 1856:279. [fuller description Engelmann 1859:31].
TL: Coahuila, between Eagle Pass & Santa Rosa, Bigelow, Jan.1853 (MO).
Synonyms:
Echinocereus dasyacanthus var. ctenoides (Engelmann) Backeberg1960:2021.
Echinocereus pectinatus var. ctenoides* Weniger 1971:31 nom. inval.
(no basionym given).
Echinocereus pectinatus var. ctenoides (Engelmann) Weniger
ex Frank 1991:135.
Echinocereus pectinatus ssp. ctenoides (Engelmann) Frank 1997:136.
Note: Echinocereus pectinatus var. wenigeri is considered by
Frank (1997) to be a synonym of this species.
Echinocereus cucumis Werdermann 1949:3-4.
TL: unknown [received from Hummel in 1937 without location noted,
but probably from Sonora, Mexico].
Synonyms:
Echinocereus noctiflorus* Hort. ex Backeberg 1960:1991 pro syn.
Echinocereus nocturniflorus* Hort. ex Backeberg 1960:1991 pro syn.
Echinocereus scheeri var. gentryi cultivar ‚Cucumis‘ (Werdermann)
Taylor 1984:155.
Echinocereus scheeri var. gentryi fa. cucumis*Parker
1996:4 nom. nud.
Note: Usually considered a synonym of Echinocereus gentryi.
Echinocereus dahliaflorus*
[An April fool prank name in Möllers Deutche Gärter-Zeitung 15:148 (1900)].
Echinocereus dasyacanthus Engelmann 1848a:100.
TL: Mexico, Chihuahua, Mtns. near Cd. Juarez, Wislizenus (MO).
Synonyms:
Cereus dasyacanthus (Engelmann) Engelmann 1849:50.
Cereus deflexispinus* Monville ex Labouret 1853:321 pro syn.
Echinocereus pectinatus var. dasyacanthus (Engelmann) Engelmann
        ex Haage 1892:118.
Echinocereus pectinatus var. dasyacanthus* Earle 1971:80
        nom. illegit. (later homonym).
Echinocereus pectinatus var. dasyacanthus* Taylor 1984:179
nom. illegit. (later homonym).
Note: Benson uses the name Echinocereus pectinatus var.
neomexicanus for this species.
Echinocereus dasyacanthus var. ctenoides (Engelmann) Backeberg 1960:2021
= Echinocereus ctenoides.
Echinocereus dasyacanthus var. hildmannii* Arendt ex Weniger
1970:33 nom. inval. (no basionym given) = Echinocereus hildmannii.
Cereus dasyacanthus var. minor* Engelmann 1856:278 nom. illegit.
  = Echinocereus roetteri.
Note: This name is rejected by the author in his addenda of corrections and
replaced by Echinocereus roetteri.
Cereus dasyacanthus var. neomexicanus Coulter 1896:384.
TL: Southeastern New Mexico,[actually Pecos County, Texas]
  Wright 366 (MO).
Synonyms:
Echinocereus pectinatus var. neomexicanus (Coulter) Benson 1944:256.
Echinocereus dasyacanthus var. neomexicanus* Coulter ex 
        Weniger 1970:32 nom. inval.
Note: Benson uses the name Echinocereus pectinatus var.
neomexicanus for Echinocereus dasyacanthus. The original var.
neomexicanus was probably a natural hybrid (see Zimmerman 1993).
Echinocereus dasyacanthus var. neomexicanus* Coulter ex 
  Weniger 1970:32 nom. inval. (no basionym given).
Echinocereus dasyacanthus var. rectispinus Trocha & Fethke 1991:31-40.
TL: Mexico, Chihuahua, near Villa Ahumada, Fiehn 1979. (Hamburg Herbarium).
Synonyms:
Echinocereus pectinatus var. rectispinus* Zimmerman 1993:277 pro syn.
Echinocereus dasyacanthus ssp. rectispinus (Trocha & Fethke)Blum, Rischer &
Rutow  inBlum et al. 1998:118.
Echinocereus dasyacanthus ssp. rectispinus (Trocha & Fethke)Blum, Rischer & Rutow
in Blum et al. 1998:118.
Echinocereus dasyacanthus var. rigidissimus (Engelmann) Marshall
& Bock 1941:119 = Echinocereus pectinatus var. rigidissimus.
Echinocereus dasyacanthus fa. roetteri* Parker 1996:2 nom. nud. 
  = Echinocereus roetteri.
Echinocereus dasyacanthus var. rubrus* Fric 1924:121 nom. nud.
Cereus dasyacanthus var. spurius Labouret 1853:321.
TL: unknown
Synonyms:
Cereus deflexispinus var. spurius  Monville ex  Labouret 1853:321.
Echinocereus dasyacanthus var. steereae (Clover) Marshall 1945:115
= Echinocereus steereae.
Echinocereus dasyacanthus fa. steereae* Parker 1996:2 nom. nud. 
  = Echinocereus steereae.
Echinocereus datae Laferriere 1997:28.
  TL: Mexico, Chihuahua, near Nabogame, Laferriere1425, 8 Apr 1988 (ARIZ).
Echinocereus davisii Houghton 1931:466.
TL: Texas, 4 mi. south of Marathon, 4000 ft., Houghton 700
(US 1566585).
Synonyms:
Echinocereus viridiflorus var. davisii (Houghton) Marshall
& Bock 1941:119.
Echinocereus viridiflorus fa. davisii (Houghton) Krainz 1967:51.
Echinocereus viridiflorus ssp. davisii (Houghton)Taylor 1997:10.
Echinocereus decumbens Clover & Jotter 1941:417.
TL: Arizona, Coconino Co., Mile 16½ Marble Canyon, Clover
& Jotter 2212 (MICH).
Synonyms:
Echinocereus engelmannii var. decumbens (Clover & Jotter)Benson 1944:258.
Echinocereus engelmannii ssp. decumbens (Clover & Jotter)
Blum & Lange in Blum et al. 1998:76.
Cereus deflexispinus* Monville ex Labouret 1853:321 pro syn. 
  = Echinocereus dasyacanthus.
Cereus deflexispinus* var. spurius Monville ex Labouret
1853:321 pro syn. 
    = Cereus dasyacanthus var. spurius.
Echinocereus degandii* Rebut ex Schumann 1895:123 nom. nud. 
  [Probably = Echinocereus dasyacanthus].
Echinocereus delaetii (Gürke) Gürke 1909b:131.
= Cephalocereus delaetii Gürke 1909a:116-121.
TL: unknown [Probably Sierra Paila, Coahuila].
Synonyms:
Echinocereus longisetus var. delaetii (Gürke) Taylor 1988:79.
Echinocereus longisetus ssp. delaetii (Gürke) Taylor 1997:9.
Echinocereus delaetii var. freudenbergeri (Frank) Taylor 1985b:97
= Echinocereus freudenbergeri.
Cereus deppei Salm-Dyck 1834:338.
TL: Mexico.
Note: This species is usually considered a synonym of Echinocereus
cinerascens.
Cereus deppei* Hort. Berol. ex Pfeiffer 1837:101 pro syn.
= Echinocereus cinerascens.
Cereus deppei* Hort. Paris ex Salm-Dyck 1845:27 nom. nud.
Echinocereus deppei *(Salm-Dyck) Schumann 1895:123 pro syn. 
  = Echinocereus cinerascens.
Echinocereus depressus* Haage 1900:157 nom. nud.
*Echinocereus diguetii* Hort. nom. nud. = Cereus diguetii.
Echinocereus dubius (Engelmann) Engelmann ex Rümpler 1885:787.
= Cereus dubius Engelmann 1856:282-283. [fuller description in Engelmann 1859:36].
TL: Texas, Hudspeth Co. near Rio Grande, Wright, 19 June 1852 (MO).
Synonyms:
Echinocereus malibrani* Rebut ex Haage 1892:118.
Echinocereus enneacanthus var. dubius (Engelmann) Benson 1969d:127.
Echinocereus enneacanthus fa. dubius* Parker 1996:2 nom. nud.
Note: This species is usually considered to be a synonym of
Echinocereus enneacanthus var. enneacanthus.
*Echinocereus dumenilianus Monville ex Haage 1859:19 = Echinopsis sp.
Echinocereus durangensis Poselger ex Rümpler 1885:799.
TL: Mexico, Durango.
Synonyms:
Echinocereus acifer var. durangensis (Poselger ex Rümpler) Schumann 1897:287.
Echinocereus polyacanthus var. durangensis * Parker 1996:4 nom. nud.
Note: Taylor (1985b) lists this name as a synonym of Echinocereus 
          polyacanthus.
Echinocereus durangensis var. nigrispinus Hort. ex Rümpler 1885:800.
TL: Mexico, Durango.
Echinocereus durangensis var. rufispinus Hort. ex Rümpler 1885:800.
TL: Mexico, Durango.
Echinocereus ehrenbergii* Engelmann ex  Haage 1859:19. nom. nud.
  [Probably = Echinocereus ehrenbergii(Pfeiffer)Pfeiffer ex Rümpler]
Echinocereus ehrenbergii (Pfeiffer)Pfeiffer ex Rümpler 1885:775.
=Cererus ehrenbergii Pfeiffer 1840:282
TL: Mexico, Hidalgo, Mineral del Monte [according to Schumann 1897:262]
     Ehrenberg (?).
Synonyms:
Echinocereus pentalophus var. ehrenbergii (Pfeiffer) Backeberg 1960:2003.
Echinocereus cinerascens var. ehrenbergii (Pfeiffer) Bravo 1974:47.
Echinocereus cinerascens ssp. ehrenbergii (Pfeiffer) Blum & Rutow in Blum et al. 1998:140-141.
*Echinocereus emoryi = Bergerocactus emoryi.
Echinocereus engelmannii (Parry ex Engelmann) Engelmann ex Haage 1859:19
= Cereus engelmannii Parry ex Engelmann 1852:328.
TL: California, Mtns. near San Felipe, Parry (?).
Echinocereus engelmannii* Lemaire 1868:56 nom. nud. 
  [Probably = Echinocereus engelmannii Parry ex Rümpler].
Echinocereus engelmannii* Harry ex  Seitz 1878:10. nom. nud.
  [Probably = Echinocereus engelmannii Parry ex Rümpler].
Echinocereus engelmannii var. acicularis Benson 1969a:22.
TL: Arizona Maricopa Co., Crossing of New River, south side of Black
Canyon Refuge, 1300 ft., Benson 16,616 20, Apr. 1966 (POM 311313).
Echinocereus engelmannii var. albispinus* Cels ex Schumann
1897:276 nom. nud.
Echinocereus engelmannii var. armatus Benson 1969c:33.
TL: Calif., San Bernardino Co., Dead Man Point, Benson 14,767,
1 Apr. 1951 (POM 284927).
Echinocereus engelmannii var. bonkerae* Mesa Garden Seed Catalog
1996 nom. nud. = Echinocereus bonkerae.
Echinocereus engelmannii var. boyce-thompsonii* Mesa Garden Seed Catalog 1996
   nom. nud. = Echinocereus boyce-thompsonii.
Echinocereus engelmannii var. caespitosus* Orcutt 1897:276 nom. nud.
Echinocereus engelmannii var. chrysocentrus (Engelmann & Bigelow)
Engelmann ex Rümpler 1885:806.
= Cereus engelmannii var. chrysocentrus Engelmann & Bigelow
in Engelmann 1856:283. [Fuller description in Englemann 1857:35]
TL: Arizona, Bill Williams River, J. M. Bigelow, 13 Feb. 1854 (MO).
Synonyms:
Echinocereus chrysocentrus (Engelmann & Bigelow) Orcutt 1926:4.
Echinocereus engelmannii var. decumbens (Clover & Jotter) Benson 1944:258
   = Echinocereus decumbens.
Echinocereus engelmannii ssp. decumbens (Clover & Jotter)Blum & Lange
in Blum et al. 1998:76.  = Echinocereus decumbens.
Echinocereus engelmannii var. fasciculatus* Mesa Garden Seed Catalog 1996
   nom. nud. = Echinocereus fasciculatus.
Echinocereus engelmannii ssp. fasciculatus(Engelmann ex Watson) Blum & Lange
inBlum et al. 1998:78. = Echinocereus fasciculatus.
Echinocereus engelmannii var. fulvispinus* Cels ex Schumann
1897:276 nom. nud.
Echinocereus engelmannii var. howei Benson 1974:80.
TL: Calif., San Bernardino Co., east of Goffs, D.F. Howe
4,570 (POM 317,886).
Echinocereus engelmannii subsp. llanuraensis ( Rutow ) Felger 1999. Proc. San Diego Soc. Nat. Hist., 35: 20
=Echinocereus nicholii subsp. llanuraensis Rutow
Echinocereus engelmannii var. munzii (Parish) Pierce & Fosberg   1933:123
= Echinocereus munzii.
Echinocereus engelmannii ssp. munzii* Blum 1997 Seed List nom. nud. 
  = Echinocereus munzii.
Echinocereus engelmannii var. nicholii Benson 1944:258.
TL: Arizona, Pima Co., Silver Bell Mtns., Benson 10,720, 28 Mar. 1941 (Ariz 34989).
Synonyms:
Echinocereus nicholii (Benson) Parfitt 1987:157-158.
Echinocereus engelmannii var. pfersdorffii* Heyder ex Schumann
1897:276 nom. nud.
Echinocereus engelmannii var. purpureus Benson 1969d:126-127.
TL: Utah, Washington Co., north of St. George, 2900 ft. Benson
13,637, 5 May 1949 (POM 285578).
Echinocereus engelmannii var. robustior* Haage 1892:117 nom. nud.
Echinocereus engelmannii var. robustior* Hildmann ex Hirscht 1894:194 nom. nud.
Echinocereus engelmannii var. variegatus (Engelmann & Bigelow)
Engelmann ex Rümpler 1885:806.
= Cereus engelmannii var. variegatus Engelmann & Bigelow
in Engelmann 1856:283. [Fuller description in Engelmann 1857:35]
TL: Arizona, Cactus Mountains at head of Bill Williams River, J.M. Bigelow,
1 Feb. 1854 (MO).
Echinocereus engelmannii ssp. variegatus* Breckwoldt in Kakteen und andere
Sukkulenten 44(8): file card 17. nom inval. (No basionym)
= Echinocereus engelmannii var. variegatus.
Echinocereus engelmannii subsp. variegatus ( Engelm. & J.M.Bigelow ) E.Lutz
Echinocereenfreund 13(4): 97. 2000
= Echinocereus engelmannii var. variegatus.
Echinocereus engelmannii var. versicolor* Hirscht 1894:194 nom. nud.
Echinocereus enneacanthus Engelmann 1848a:111.
TL: Mexico, Chihuahua, near San Pablo, Wislizenus 244, 8 Apr.
1847 (MO 83797).
Synonyms:
Cereus enneacanthus (Engelmann) Engelmann 1849:50.
Note: Echinocereus lowryi* Lowry 1936:20 nom. nud. and Echinocereus
         albiflorus* Lowry 1937:20 nom. nud. are probably forms of this species.
Echinocereus enneacanthus fa. brevispinus Moore 1967:93-94.
TL: Texas, Starr Co., 8 miles East of Rio Grande City, Clover, 31
Dec 1953 (MICH).
Synonyms:
Echinocereus enneacanthus var. brevispinus (Moore) Benson
1969d:127
      Echinocereus enneacanthus ssp. brevispinus (Moore) Taylor 1997:8.
Echinocereus enneacanthus var. brevispinus (Moore) Benson 1969d:127
= Echinocereus. enneacanthus fa. brevispinus.
Echinocereus enneacanthus ssp. brevispinus (Moore) Taylor 1997:8.
  = Echinocereus enneacanthus fa. brevispinus.
Echinocereus enneacanthus var. brevispinus fa. carnosus* Parker 1996:2 nom. nud.
    = Echinocereus carnosus.
Echinocereus enneacanthus var. carnosus (Rümpler) Schelle1907:127
    = Echinocereus carnosus.
Echinocereus enneacanthus var. carnosus* Quehl 1908:114 nom. nud. 
  [Probably = Echinocereus carnosus].
Echinocereus enneacanthus fa. carnosus* 
  is attributed incorrectly to Matthson ex Schumann by Bravo 1991:76.
Echinocereus enneacanthus var. conglomeratus (Blanc) Benson 1974:80
= Echinocereus conglomeratus.
Echinocereus enneacanthus var. dubius (Engelmann) Benson 1969d:127
= Echinocereus dubius.
Echinocereus enneacanthus fa. dubius* Parker 1996:2 nom. nud. 
  = Echinocereus dubius.
Echinocereus enneacanthus var. erectus* Haage 1892:117 nom. nud.
Echinocereus enneacanthus fa. intermedius Moore 1967:93.
TL: Texas, Starr Co., off La Grulla Road, Clover, 23 Mar. 1953
(MICH).
Note: Usually considered a synonym of Echinocereus enneacanthus 
    var. brevispinus.
Echinocereus enneacanthus var. major* Schelle 1907:127 nom. nud.
Echinocereus enneacanthus var. majus* Haage 1892:117 nom. nud
Echinocereus enneacanthus fa. merkeri* Parker 1996:2 nom. nud. 
  = Echinocereus merkeri.
Echinocereus enneacanthus fa. sarissophorus* Parker 1996:2 nom. nud. 
  = Echinocereus sarissophorus..
Echinocereus enneacanthus var. stramineus (Engelmann) Benson
1969d:127    = Echinocereus stramineus.
Echinocereus enneacanthus fa. uspenskii* Parker 1996:2 nom. nud. 
  = Echinocereus uspenskii.
*Echinocereus erectocentrus var. palliduserror in spelling = Echinomastus erectocentrus.
Echinocereus fasciculatus (Engelmann) Benson 1969a:21.
= Mammillaria fasciculata* Engelmann in Emory 1848:156. nom. prov.
TL: Arizona, Graham Co. 6-7 miles east of Solomon. [Neotype –
Benson 16633, 22 April 1966 (POM 311339)].
Synonyms:
Echinocereus fendleri var. fasciculatus (Engelmann) Taylor
1985a:252.
Echinocereus engelmannii var. fasciculatus* Mesa Garden
Seed Catalog 1996 nom. nud.
        Echinocereus engelmannii ssp. fasciculatus (Engelmann ex Watson) Blum &
Lange in Blum et al. 1998:78
Note: The validity of this name is uncertain. Echinocereus robustus is usually
considered to be a synonym of this plant.The name Mammillaria fasciculata is
listed as a synonym of Cereus fendleri by Watson 1878:404.
Echinocereus fasciculatus ssp. abbeae* Blum 1997 Seed List nom. nud. 
   = Echinocereus abbeae.
Echinocereus fasciculatus var. bonkerae (Thornber & Bonker) Benson 1969a:22
   = Echinocereus bonkerae.
Echinocereus fasciculatus ssp. bonkerae  (Thornber & Bonker) Taylor 1997:8
   = Echinocereus bonkerae.
Echinocereus fasciculatus var. boyce-thompsonii (Orcutt) Benson 1969a:21
= Echinocereus boyce- thompsonii.
Echinocereus fasciculatus ssp. boyce-thompsonii (Orcutt) Taylor 1997:8
    = Echinocereus boyce-thompsonii.
Echinocereus fasciculatus var. rectispinus* Benson 1969a:131 pro syn.
  = Echinocereus rectispinus.
Echinocereus fasciculatusfa. robustus* Parker 1996:2 nom. nud. 
  = Echinocereus robustus.
Echinocereus fendleri* Sencke ex  Seitz 1870:11 nom. nud. 
    [Probably a synonym of Echinocereus fendleri Loder].
Echinocereus fendleri (Engelmann) Loder 1882:445.
= Cereus fendleri Engelmann 1849:51.
TL: New Mexico, near Santa Fe, Fendler 3, 1846 (MO).
Synonyms:
Echinocereus melaleuca* Föster ex Haage 1892:117 nom. nud.
Cereus cinerascens var. fendleri (Engelmann) Bois 1928:371.
Note: Usually listed as Echinocereus fendleri Engelmann ex
Rümpler 1885:801 or Senke ex Haage. 1893:22.
Echinocereus fendleri var. albiflorus (Weingart) Backeberg 1960:2047
  = Echinocereus albiflorus.
Echinocereus fendleri fa. albiflorus* Parker 1996:2 nom. nud. 
  = Echinocereus albiflorus Weingart.
Echinocereus fendleri var. bonkerae (Thornber & Bonker) Benson 1944:260
= Echinocereus bonkerae.
Echinocereus fendleri var. boyce-thompsonii (Orcutt) Benson 1944:260
= Echinocereus boyce-thompsonii.
Echinocereus fendleri var. fasciculatus (Engelmann) Taylor 1985a:252
= Echinocereus fasciculatus.
Echinocereus fendleri var. hempelii* Mesa Garden Seed Catalog
1996 nom. nud.     = Echinocereus hempelii
Echinocereus fendleri ssp. hempelii Blum in Blum et al. 1998:93.
= Echinocereus hempelii.
Echinocereus fendleri var. kuenzleri (Castetter et al.) Benson 1982:942
= Echinocereus kuenzleri.
Echinocereus fendleri ssp. kuenzleri* Blum 1997 Seed List nom. nud. 
  = Echinocereus kuenzleri.
Echinocereus fendleri var. ledingii (Peebles) Taylor 1985a:253
= Echinocereus ledingii.
Echinocereus fendleri var. major* Haage 1900:158 nom. nud.
Echinocereus fendleri var. pauperculus (Engelmann) Schumann 1897:275.
= Cereus fendleri var. pauperculus Engelmann 1849:51.
TL: New Mexico, near Santa Fe, Fendler, 4 Nov. 1846 (MO).
Echinocereus fendleri var. rectispinus (Peebles) Benson 1944:259
= Echinocereus rectispinus.
Echinocereus fendleri ssp. rectispinus (Peebles) Taylor 1997:9
  = Echinocereus rectispinus.
Echinocereus fendleri var. robustus Fobe 1911:55
[not Benson 1944].
TL: unknown
Echinocereus fendleri var. robustus* (Peebles) Benson 1944:259
  nom. illegit (later homonym).    = Echinocereus robustus.
Echinocereus ferreirianus Gates 1953:8-11.
TL: Mexico, Baja Calif. Norte, Isla Piojo, Bahia de los Angeles,
H.E. Gates, 1934 (DS 375348).
Echinocereus ferreirianus var. lindsayi (Meyran) Taylor 1985b:46
= Echinocereus lindsayi.
Echinocereus ferreirianus ssp. lindsayi (Meyran) Taylor 1997:9
= Echinocereus lindsayi.
Echinocereus ferreirrae* Gates 1935:154 nom. nud. 
(= Gates number 550).
Echinocereus fiehnii* Lau 1992:74 nom. nud. = Echinocereus russanthus var. fiehnii.
Echinocereus finnii* Hort. ex Taylor 1985 101 pro syn.
= Echinocereus russanthus var. fiehnii.
Echinocereus fitchii Britton & Rose 1922:30.
TL: Texas, near Laredo, Rose 18037, 1913 (US).
Synonyms:
Echinocereus reichenbachii var. fitchii (Britton & Rose) Benson 1969d:127.
Echinocereus reichenbachii ssp. fitchii (Britton & Rose) Taylor 1997:9.
Echinocereus fitchii var. albertii* Mesa Garden Seed Catalog 996 nom. nud.
   = Echinocereus reichenbachii var. albertii.
Echinocereus fitchii ssp. albertii (Benson)Blum & Lange in Blum et al. 1998:313.
    = Echinocereus reichenbachii var. albertii.
Echinocereus fitchii var. armatus* Mesa Garden Seed Catalog1996 nom. nud.
    = Echinocereus pectinatus var. armatus.
Echinocereus fitchii ssp. armatus (Poselger) Blum, Lange & Rutow in
Blum et al. 1998:314.
    = Echinocereus pectinatus var. armatus.
*Echinocereus flavescens   = Haageocereus sp.
Echinocereus flavidiflorus* Haage 1892:117 nom. nud.
Echinocereus flavidiflorus* Hildmann ex Schumann 1897:264 pro syn. 
  = Echinocereus leonensis.
Echinocereus flaviflorus (Coulter) Coulter ex Schumann 1897:274.
= Cereus flavidiflorus Engelmann ex Coulter 1896:391.
TL: Mexico, Baja California, west of San Borgia, Gabb 10,
1867(MO).
Note: Usually considered a synonym of Echinocereus maritimus.
Echinocereus flavispinus* Bailey 1941:270 nom. nud.
Echinocereus floresii Backeberg 1949:5.
TL: Mexico, Sinaloa, near Topolobampo, R. Flores & F. Schwarz (?).
Synonyms:
Echinocereus sciurus var. floresii (Backeberg) Taylor1985b:115.
Echinocereus sciurus ssp. floresii (Backeberg) Taylor1997:10.
Echinocereus bristolii ssp. floresii Blum & Lange in Blum et al. 1998:353.
Echinocereus fobeanus Oehme 1940:49-50.
TL: unknown [Probably Coahuila].
Synonyms:
Echinocereus chisoensis var. fobeanus (Oehme) Taylor 1985a:261.
Note: Taylor (1993) lists Echinocereus metornii as a synonym
of this species.
Echinocereus fobeanus ssp. metornii (Frank) Blum & Lange in Blum et al. 1998:307.
    = Echinocereus metornii
Echinocereus fordtii* spelling error for  Echinocactus fordtii.
Echinocereus freudenbergeri Frank 1981:102-105.
TL: Mexico, Coahuila, 12 km south of Cuatro Cienegas, G. 
  Freudenberger (ZSS AA18-27).
Synonyms:
Echinocereus delaetii var. freudenbergeri (Frank) Taylor
1985b:97.
Echinocereus setosus* Hort. ex Taylor 1988:76 pro syn.
Echinocereus nivosus var. freudenbergeri* Parker 1996:3  nom. nud.
      Echinocereus longisetus ssp. freudenbergeri (Frank) Blum in Blum et al. 1998:187.
Note: Taylor (1993) lists this species as a synonym of
Echinocereus nivosus.
Echinocereus galtieri* Hort. Grus. ex Schumann 1895:124 nom. nud.
Echinocereus galtieri Rebut ex Fobe 1911:56.
TL: unknown
Synonyms:
Echinocereus polyacanthus var. galtieri (Rebut ex Fobe) Schelle 1926:179.
Echinocereus gentryi Clover 1938:565.
TL: Mexico, Sonora, Cañon Saucito, H.S. Gentry, 1932 (MICH).
Synonyms:
Echinocereus scheeri var. gentryi (Clover) Taylor 1984:154.
Echinocereus scheeri ssp. gentryi (Clover) Taylor 1997:9
Note: Echinocereus cucumis is usually considered a
synonym of this species.
*Echinocereus gladiatus 
  = Echinopsis gladiata.
Echinocereus glycerius* Förster ex Haage 1859:19 nom. nud.
Echinocereus glycimorphus* Förster ex Seitz 1870:11 nom. nud.
Echinocereus glycimorphus Förster ex Rümpler 1885:800.
TL: unknown [Hidalgo near Ixmiquilpan and El Cardonal according
to Schumann 1897:262].
Synonyms:
Cereus glycimorphus (Förster ex Rümpler) Orcutt 1903:5.
Echinocereus cinerascens fa. glycimorphus* Parker 1996:2 nom. nud. 
Echinocereus glycimorphus var. undulatus Schelle 1926:166.
TL: unknown
Note: Schelle 1926:166 lists Echinocereus undulatus* Hildmann
as a synonym of this variety.
Cereus glomeratus* Orcutt 1897:21 pro syn.= Echinocereus maritimus.
Echinocereus gonacanthus* Lemaire 1868:57 nom. nud.
[Probably = Echinocereus gonacanthus Loder].
Echinocereus gonacanthus (Engelmann & Bigelow) Loder 1882:444.
= Cereus gonacanthus Engelmann & Bigelow in Engelmann 1856:283.
[fuller description Engelmann 1857:33].
TL: Arizona, Apache Co., Jacobs Well, 40 mi. west of Zuñi,
    J. M. Bigelow, 29 Nov. 1853 (MO).
Synonyms:
Echinocereus paucispinus var. gonacanthus (Engelmann &
Bigelow) Schumann 1897:281.
Echinocereus triglochidiatus var. gonacanthus (Engelmann
& Bigelow) Boissevain 1940:36.
Note: Usually listed as Echinocereus gonacanthus
Engelmann ex Rümpler 1885:806.
*Echinocereus grahamii (Monatschrift Kakteenkunde 20:47)
error for Mammillaria grahamii.
Echinocereus grandis Britton & Rose 1922:18.
TL: Mexico, Gulf of California, San Esteban Island, J.N. Rose
16823, 13 Apr. 1911 (US).
Synonyms:
Echinocereus pectinatus var. grandis (Britton & Rose) Marshall 1955:68.
Echinocereus hancockii Dawson 1949:89-93.
TL: Mexico, Baja California Sur, north sideof San Hipolito
Bay, E. Y. Dawson 6443, 4 April 1949(LAM).
Synonyms:
Echinocereus maritimus var. hancockii (Dawson) Taylor 1985b:44.
Echinocereus maritimus ssp. hancockii (Dawson) Blum & Rutow in
Blum et al. 1998:57.
Echinocereus havermansii* Rebut ex Haage 1892:118 nom. nud.
Echinocereus havernicki* Hildm. ex Schwarzbach & Bitter1915:8 nom. nud.
Echinocereus hempelii Fobe 1897:187.
TL: Mexico [Probably Santa Clara Canyon, Chihuahua, Mexico].
Synonyms:
Echinocereus fendleri var. hempelii * Mesa Garden Seed Catalog 1996 nom. nud.
      Echinocereus fendleri ssp. hempelii Blum in Blum et al. 1998:93.
Note: Some authors treat Echinocereus kuenzleri as a
synonym of this species.
Echinocereus hempelii var. centrispinus* Hort. nom. nud.
[=HK 1489 from San Buenaventura, Chihuahua, Mexico]
Echinocereus hexaedrus (Engelmann & Bigelow) Engelmann ex 
  Rümpler 1885:807.
= Cereus hexaedrus Engelmann & BigelowEngelmann 1856:285.
[fuller description Engelmann 1857:34].
TL: Arizona, Apache County, 15 miles west of Zuñi, J. M. 
      Bigelow, 28 Nov. 1853 (MO).
Synonyms:
Echinocereus paucispinus var. hexaedrus (Engelmann &
Bigelow) Schumann 1897:281.
Echinocereustriglochidiatus var. hexaedrus (Engelmann
& Bigelow) Boissevain 1940:36.
Echinocereuscoccineus fa. hexaedrus* Parker 1996:4 nom. nud.
Note: Considered by Taylor (1985b) as a synonym
of Echinocereus triglochidiatus  var.  melanacanthus.
Echinocereus hidalgensis* Hort. Berol. ex Backeberg 1962:3855-3856 nom. nud.
Echinocereus hildmannii Arendt 1892:146.
TL: unknown [West Texas according toSchelle 1926:175].
Synonyms:
Echinocereus dasyacanthus var. hildmannii* Arendt ex Weniger
1970:33 nom. inval.
Note: This plant may be a natural hybrid (see Zimmermann 1993).
Echinocereus huitcholensis (Weber) Gürke 1906:23.
= Cereus huitcholnesis Weber 1904:383.
TL: Mexico, Jalisco, Sierra de los Huitcholes, 1800 m, L. 
    Diguet, May 1900 (P).
Synonyms:
Echinocereus polyacanthus var. huitcholensis (Weber) Taylor1988:82.
Echinocereus acifer ssp. huitcholensis (Weber) Lange 1994:109.
Echinocereus polyacanthus var. acifer fa. huitcholensis*Parker 1996:4 nom. nud.
        Echinocereus polyacanthus ssp. huitcholensis (Weber) Taylor 1997:9.
*Echinocereus hypogaeus  = Eridisia spiniflora.
Echinocereus inermis* Haage ex Hirscht 1898:130 pro syn.
= Echinocereus knippelianus.
*Echinocereus intricatus = Echinopsis strigosa.
Echinocereus jacobyi* Hort. ex Schumann 1897:278 pro syn.
= Echinocereus merkeri.
Echinocereus joconosle* Haage 1892:117 nom. nud.
Echinocereus klapperi Blum in Blum et al. 1998:380-383.
TL: Mexico, Sonora, El Novillo,500 m, Klapper 0121, 12 March 1983
(ZSS A 17985).
Echinocereus knippelianus Liebner 1895:170.
TL: unknown [Relocated in Sierra Parras, Coahuila, Mexico].
Synonyms:
Echinocereus liebnerianus* Liebner ex Schumann 1896b:262
        nom. nud.
Echinocereus inermis* Haage ex Hirscht 1898:130 pro syn.
Cereus knippelianus (Liebner) Orcutt 1902:27.
Echinocereus knippelianus var. kruegeri Glass & Foster 1978b:80.
TL: Mexico, Nuevo Leon, north of Ascencion near San Juanito,
  Glass & Foster 3902, 1 Mar. 1972 (POM).
Echinocereus knippelianus ssp. kruegeri (Glass & Foster) Blum & Lange in Blum et al. 1998:248. = Echinocereus knippelianus var. kruegeri.
Echinocereus knippelianus var. kruegeri fa. reyesii* Parker 1996:3 nom. nud. 
    = Echinocereus knippelianus var. reyesii.
Echinocereus knippelianus var. reyesii Lau 1980:264-265.
TL: Mexico, Nuevo Leon, between Siberia & Encantada, Lau 1237A,
Apr. 1977 (POM).
Synonyms:
Echinocereus knippelianus var. kruegeri fa. reyesii*Parker 1996:3 nom. nud.
Echinocereus knippelianus ssp. reyesii (Lau) Blum & Lange in Blum et al.1998:250.
= Echinocereus knippelianus var. reyesii.
Echinocereus knippelianus var. schaetzleii* Nitzschke 1990:43 nom.  prov.
Echinocereus knippelianus var. siberi giberia* Heyer 1989:26 nom. nud.
Echinocereus koehresianus* Hort. ex Taylor 1988:80 pro syn.
= Echinocereus scheeri var. koehresianus.
Echinocereus koehresii* Hort. ex Taylor 1988:80 pro syn. 
  = Echinocereus scheeri var. koehresianus.
Echinocereus krausei De Smet ex Rümpler 1885:789.
TL: unknown
Note: This plant is usually listed as a synonym of Echinocereus coccineus.
Echinocereus kroenleinii* Hort. nom. nud = Wilcoxia kroenleinii
Wilcoxia kroenleinii Cartier 1980 (Succulentes 2(2):2-3).
TL: unknown, HK 379 (P) [Probably from Coahuila, Mexico]
Synonyms:
Echinocereus poselgeri ssp. kroenleinii (Cartier) Lange 1995:138.
Echinocereus poselgeri fa. kroenleinii* Parker 1996:6  nom. nud.
Echinocereus kroenleinii* Hort.
Echinocereus kuenzleri Castetter et al. 1976:77-78.
TL: New Mexico, Otero Co., Sacramento Mtns., Elk, Künzler 3585,
5 May 1968 (UNM 55571).
Synonyms:
Echinocereus pseudohempelii* Künzler Cat. nom. nud.
Echinocereus fendleri var. kuenzleri (Castetter et al.) Benson 1982:942.
Echinocereus fendleri var. hempelii fa. kuenzleri* Parker 1996:2 nom. nud.
Echinocereus fendleri ssp. kuenzleri* Blum 1997 Seed List nom. nud.
Echinocereus kunzei Gürke 1907:103-104.
TL: Arizona, near Phoenix[southern New Mexico according to
Britton & Rose 1922:31].
Synonyms:
Echinocereus coccineus var. kunzei (Gürke) Backeberg
1960:2070.
Note: This plant may be a natural hybrid. (see Zimmer-
mann 1993).
Echinocereus laboretianus Daul 1890:76.
TL: unknown
Note:Usually considered a synonym of Echinocereus viridiflorus.
Echinocereus labouretianus* Lemaire 1868:57 nom. nud.
[Probably = Echinocereus laboretianus]
Echinocereus labouretianus var. chrysacanthus* Haage 1900:159 nom. nud.
Echinocereus labouretianus var. gracilispinus* Haage 1900:159 nom. nud.
Echinocereus labouretianus var. sanguineus* Haage 1900:159 nom. nud.
Echinocereus labouretii* Förster ex Haage 1859:19  nom. nud.
Echinocereus labouretii* Förster ex Rümpler 1885:811 pro syn. 
  = Echinocereus viridiflorus.
*Echinocereus lamprochlorus = Echinopsis lamprochlora.
Echinocereus laui Frank 1978:74-77.
TL: Mexico, Sonora, east of Yecora, El Trigo Ranch, 1800m,
A. Lau 780, Jan. 1974 (ZSS AA18- 24).
Echinocereus ledingii Peebles 1936:35.
TL: Arizona, Graham Co., Mt. Graham, 4500 ft., Louis Wankum, 11
July 1935 (US 1634004).
Synonyms:
Echinocereus fendleri var. ledingii (Peebles) Taylor 1985a:253.
Echinocereus leeanus* Lemaire 1868:57 nom. nud. 
[Probably = Echinocereus leeanus Lemaire ex Rümpler].
Echinocereus leeanus (Hooker) Lemaire ex Rümpler 1885:828.
= Cereus leeanus Hooker 1849:t4417.
TL: Northern Mexico.
Synonyms:
Echinocereus polyacanthus var. leeanus* Parker 1996:4 nom. nud. 
Note: Often considered a synonym of Echinocereus polyacanthus.
Echinocereus leeanus var. multicostatus (Cels ex Rümpler) Schumann 1897:289
= Echinocereus multicostatus.
Echinocereus leonensis Mathsson 1891:66.
TL: Mexico, Nuevo Leon, near Monterrey, 1800ft. Runge (?).
Synonyms:
Echinocereus flaviflorus* Hildmann ex Schumann 1897:264  pro syn.
Cereus leonensis (Mathsson) Berger 1905:80.
Echinocereus blanckii var. leonensis (Mathsson) Backeberg1960:1999.
Echinocereus pentalophus var. leonensis (Mathsson)Taylor 1985b:78.
Echinocereus pentalophus ssp. leonensis (Mathsson) Taylor 1997:9.
Cereus leptacanthus* De Candolle ex Pfeiffer 1837:101 pro syn. 
  = Echinocereus pentalophus var. subarticulatus.
Echinocereus leptacanthus Gaillard 1864:209
= Cereus leptacanthus De Candolle ex Salm Dyck 1845:27. which is
new namefor Cereus pentalophus var. subarticulatus De Candolle.
Cereus leptacanthus var. crassior* De Candolle ex Förster
1846:373 nom. nud.
Echinocereus leptophus* Gässner Hauptverzeichnis 1937:6 nom. nud.
Echinocereus leucanthus Taylor 1985b:136-137
[New name needed because Echinocereus albiflorus already used
for another plant]
= Wilcoxia albiflora Backeberg 1952:16.
TL: unknown [Probably northwest Sinaloa, Mexico]. (P)
[cultivated material from Les Cedres]
Echinocereus leucodictius* Engelmann ex Haage 1859:19 nom. nud.
Echinocereus liebnerianus* Liebner ex Schumann 1896b:262 nom. nud. 
  [Probably a printing error] = Echinocereus knippelianus.
*Echinocereus limensis  = Haageocereus sp .
Echinocereus lindsayi Meyran 1975:80-83.
TL: Mexico, Baja Calif. Norte, between Cataviña & Laguna
Chapala, Meyran et al., 8 May 1975 in H. Sanchez-Mejorada 2424 (MEXU)
Synonyms:
Echinocereus ferreirianus var. lindsayi (Meyran) Taylor 1985b:46.
Echinocereus ferreirianus ssp. lindsayi (Meyran) Taylor 1997:9
Echinocereus llanuraensis ( Rutow ) W.Blum & D.Felix Echinocereenfreund 15(1): 16 (2002).
= Echinocereus nicholii (L.D.Benson) B.D.Parfitt subsp. llanuraensis Rutow
Echinocereus lloydii Britton & Rose 1922:37-38.
TL: Texas, Pecos Co.,Tuna Springs, F.E. Lloyd, 26-28
Feb. 1909 (US 691964).
Synonyms:
Echinocereus roetteri var. lloydii (Britton & Rose) Backeberg 1960:2027.
Echinocereus coccineus fa. lloydii* Parker 1996:4 nom. nud.
Note: This species is probably a natural hybrid.
Echinocereus longisetus* Lemaire 1868:57 nom. nud.
[Probably = Echinocereus longisetus Engelmann ex Rümpler].
Echinocereus longisetus (Engelmann) Engelmann ex Rümpler 1885: 822.
= Cereus longisetus Engelmann 1856:280. [fuller description
Engelmann 1859:32-33, t. 45].
TL: Mexico, Coahuila, Mtns. near Santa Rosa [Melchor Muzquiz],
J.M. Bigelow, 1853 (MO).
Echinocereus longisetus var. albatus* Sterk 1980:12-13 nom. inval.
  = Echinocereus albatus* Backeberg nom. inval.
Echinocereus longisetus var. delaetii (Gürke) Taylor 1988:79
= Echinocereus delaetii.
Echinocereus longisetus ssp. delaetii (Gürke) Taylor 1997:9 = Echinocereus delaetii.
Echinocereus longisetus ssp. freudenbergeri (Frank)Blum in
Blum et al. 1998:187.= Echinocereusfreudenbergeri
Echinocereus longispinus* Hort. ex Schelle 1926:180 pro syn.
= Echinocereus acifer.
Echinocereus longispinus Lahman 1950:128.
[Originally published Lahman 1936:15 withoutLatin diagnosis].
TL: Oklahoma, Mt. Scott, Lahman (MO).
Note: Usually considered a synonym of Echinocereus baileyi.
Echinocereus longispinus var. albispinus* Backeberg 1960:2010 pro syn.
= Echinocereus albispinus Lahman.
Echinocereus lowryi* Lowry 1936:20 nom. nud. 
[Probably a form of Echinocereus enneacanthus].
Echinocereus luteus Britton & Rose 1913:239.
TL: Mexico, Sonora, Mtns. above Alamos, Rose et al. 15207, 19 March 1910
(US 535975).
Synonyms:
Echinocereus subinermis var. luteus (Britton & Rose) Knuth in
Backeberg & Knuth 1935:312.
Echinocereus subinermis fa. luteus* Parker 1996:4 nom. nud.
Echinocereus madrensis Patoni 1910:42.
TL: Mexico, Durango, high Sierra Madre.
Synonyms:
Echinocereus adustus var. schwarzii fa. madrensis* Parker 1996:6. nom. nud.
Echinocereus malibrani* Rebut ex Haage 1892:118 nom. nud.
[Probably = Echinocereus dubius (According toSchelle 1907:135)].
Echinocereus mamillatus (Engelmann ex Coulter) Britton &
Rose 1922:41.
= Cereus mamillatus Engelmann ex Coulter 1896:405.
TL: Mexico, Baja California, south of Mulege, Gabb 16,1867 (MO).
Synonyms:
Echinocereusbrandegeei fa. mamillatus* Parker 1996:1 nom. nud.
Note: Usually considered a synonym of Echinocereus brandegeei.
Echinocereus mamillifer* Förster ex Haage 1859:19 nom. nud.
Echinocereus mamilliter* Förster ex Seitz 1870:11 nom. nud.
Echinocereus mamillosus Hort. ex Rümpler 1885:787
TL: unknown
Echinocereus mapimiensis Anderson et al 1998:284-285.
TL: Mexico, Coahuila, Bolson de Mapimi, Keil 8136, 22 Aug 1971 (DES)
Echinocereus marginii* Ely 1935:6 nom. nud.
Echinocereus mariae* Backeberg 1965:2, 19 nom. inval. 
(no type designated).
Note: A form of Echinocereus baileyi from northwest of Granite,
Oklahoma.
Echinocereus maritimus (Jones) Schumann 1897:273-274.
= Cereus maritimus Jones 1883:973.
TL: Mexico, Baja California, Norte, near Ensenada, M.E. Jones,
    April 1882 (POM).
Synonyms:
Cereus glomeratus* Orcutt 1897:21 pro syn.
Note: Echinocereus orcuttii and Echinocereus flaviflorus
Coulter ex Schumann are usually considered synonyms of this species.
Echinocereus maritimus var. hancockii Taylor 1985b:44 = Echinocereus hancockii.
Echinocereus maritimus ssp. hancockii (Dawson) Blum & Rutow in
Blum et al. 1998:57.= Echinocereus hancockii.
Echinocereus marksianus* Schwarz ex Backeberg 1966:124 nom. inval. 
  (no type designated).
Note: From Northern Mexico. Taylor (1985b) lists this name as
a synonym of Echinocereus polyacanthus var. densus.
Echinocereus matlesianus*Backeberg ex Bravo 1991:26
misspelling = Echinocereus matthesianus*Backeberg.
Echinocereus matthesianus*Backeberg 1963:6 nom. inval. (no type designated).
Note: Taylor (1993) lists this name as a synonym of Echinocereus polyacanthus.
Echinocereus matudae Bravo 1961:119-121.
TL: Mexico, Chihuhaua, Rio Papigochic near Temosochic, Matuda
(MEXU).
Synonyms:
Echinocereus arizonicus ssp. matudae (Bravo) Rutow in Lange
& Rutow 1994:51.
Echinocereus coccineus var. arizonicus fa. matudae* Parker
1996:4 nom. nud.
Echinocereus melaleuca* Föster ex Haage 1892:117 nom. nud.
[Probably= Echinocereus fendleri].
Echinocereus melanacanthus* Engelmann ex Earl 1963:61
printing error.= Echinocereus coccineus.
Echinocereus melanocentrus* Lowry 1936:20 nom. inval. 
(no Latin diagnosis) = Echinocereus melanocentrus* Lowry ex Backeberg.
Echinocereus melanocentrus* Lowry ex Backeberg 1960:2030 nom. inval. 
  (no type designated).
Note: usually considered a synonym of Echinocereus reichenbachii var.albertii.
Echinocereus merkeri* Hildmann ex Haage 1892:118 nom. nud.
Echinocereus merkeri Hildmann ex Schumann 1897:277.
TL: unknown[range given as Durango near Cd. Lerdo, Coahuila,
Sierra Bola & near Parras].
Synonyms:
Echinocereus jacoybi* Hort. ex Schumann 1897:278 pro syn.
Cereus merkeri (Hildmann ex Schumann) Berger 1905:81.
Echinocereus enneacanthus fa. merkeri* Parker 1996:2 nom. nud. 
Note: Taylor (1985b) lists this species as a synonym of
Echinocereus enneacanthus.
Echinocereus merkeri var. densispinus* Schmoll ex Backeberg
1960:2051 nom. nud.
Echinocereus metornii Frank 1990b:210-218.
TL: Mexico, Coahuila, Sierra Mojada, 1500m, Metorn 49, 4 April
1989 (ZSS).
Synonyms:
Echinocereus chisoensis var. metornii* Parker 1996:5 nom. nud.
Echinocereus fobeanus ssp. metornii (Frank)Blum & Lange in Blum et al. 1998:307.
Note: Taylor (1993) lists this species as a synonym of Echinocereus fobeanus.
Echinocereus milleri Blum, Künzler & Oldach. in Blum & Oldach1999:69-76.
TL: Robert Lee, Coke Co., Texas.  Leuck 229, August 1978 (OKL).
Synonyms:
    Echinocereus russanthus ssp. milleri* Blum 1997 Seed List nom. nud
Echinocereus moelleri* Hort. (Cactus & Succulents Journal
(US) 13:22 1941) nom. nud.
Echinocereus mojavensis (Engelmann & Bigelow) Engelmann ex 
  Rümpler 1885:803.
= Cereus mojavensis Engelmann & Bigelow in Engelmann
1856:281.[Fuller description Engelmann 1857:33].
TL: California, Mojave Creek, J.M. Bigelow, 4 March 1854 (MO).
Synonyms:
Cereus bigelovii* Engelmann 1856:pl. 4 nom. nud.
Echinocereus triglochidiatus var. mojavensis (Engelmann &
Bigelow) Benson 1944:255.
Echinocereus coccineus var. mojavensis* Parker 1996:4  nom. nud.
          Echinocereus coccineus ssp. mojavensis (Engelmann & Bigelow) Taylor 1997:8.
Echinocereus triglochidiatus ssp. mojavensis (Engelmann &
Bigelow) Blum & Lange in Blum et al. 1998:362.
Echinocereus mojavensis var. albispinus* Hort. ex Schelle1926:167 nom. nud.
Echinocereus mojavensis fa. inermis* Mesa Garden Seed Catalog1996 nom. nud. 
    = Echinocereus coccineus var. inermis.
Echinocereus mojavensis var. zuniensis (Engelmann & Bigelow)
Engelmann ex Rümpler 1885:803.
= Cereus mojavensis var. zuniensis Engelmann & Bigelow in
Engelmann 1856:281.
TL: Arizona, Cañon Diablo, on Little Colorado River, J.M.
Bigelow,14 Dec. 1853 (MO).
Synonyms:
Cereusbigelovii var. zuniensis* Engelmann 1857: t. 4 nom. nud.
Echinocereus zuniensis* Eng. & Big. ex Haage & Schmidt nom. nud.
Echinocereus mombergerianus Frank 1990c:261
(Originally published Frank 1989:272-277 with no type designated).
TL: Mexico, Baja California Norte, Sierra San Pedro Martir,
Hidalgo Observatory, Lau 1248 (ZSS).
Synonyms:
Echinocereus polyacanthus var. mombergerianus* Parker1996:4 nom. nud.
Echinocereus polyacanthus ssp. mombergerianus (Frank)
Breckwoldt & Matylewicz 1996:15.
Echinocereus pacificus ssp. mombergerianus Blum, Rischer &
Rutow in Blum et al. 1998:418.
Echinocereus monacanthus Heese 1904:215.
TL: Texas – Mexico Border, Rancho Tacubaya.
Note: Usually considered a synonym of Echinocereus paucispinus.
Las Cruces, New Mexico 1 May 2006. Permission to copy or reprint
for noncommercial purposes is granted.

Echinocereus morricalii – Echinocereus reichenbachianus

Echinocereus morricalii Riha 1975:75.
TL: Mexico, Nuevo Leon, Barranca de las Garrapatassouth of
Monterrey, D. B. Morrical (PRC 322).
Synonyms:
Echinocereus viereckii var. morricalii (Riha) Taylor 1985b:93.
Echinocereus blanckii var. inermis* Heyer 1989:26 pro syn.
Echinocereus viereckii ssp. morricalii (Riha) Taylor 1997:10.
*Echinocereus multiangularis = Haageocereus sp.
*Echinocereus mutitangularis var. limensis = Haageocereus sp.
*Echinocereus multiangularis var. maletianus = ?
*Echinocereus multiangularis var. pallidior  = ?
Echinocereus multicostatus Cels ex Rümpler 1885:834.
TL: unknown
Synonyms:
Echinocereus leeanus var. multicostatus (Cels ex Rümpler)
Schumann 1897:289.
Echinocereus triglochidiatus var. multicostatus (Cels ex
Rümpler) Marshall 1953:67.
Note: this plant is often considered a synonym of
          Echinocereus leeanus, but its may be of South American
origin and not an Echinocereus.
Echinocereus munzii (Parish) Benson 1941:361.
= Cereus munzii Parish 1926:48.
TL: California, Riverside Co., 2 miles below Kenworthy, 1400m,
Munz & Johnson 5570, 21 May 1922(UC).
Synonyms:
Echinocereus engelmannii var. munzii (Parish) Pierce & Fosberg 1933:123.
Echinocereus neomexicanus Standley 1908:87-88.
TL: New Mexico, Doña Ana Co., mesa west of Organ Mtns.,
Standley 383, 1907(MO).
Synonyms:
Echinocereus triglochidiatus var. neomexicanus (Standley)
Standley ex Marshall & Bock 1941:118.
Cereus neomexicanus (Standley) Tidestrom 1941:298
Echinocereus polyacanthus var. neomexicanus* Weniger1970:44 nom. inval.
Echinocereus coccineus var. neomexicanus* Parker 1996:4 nom. nud.
Note:This plant has been confused with Echinocereus rosei
          under the name Echinocereus triglochidiatus var.  neomexicanus [Benson] and
         Echinocereus polyacanthus  [Bravo 1991:24].It is probably a natural hybrid of E. 
          rosei and E. chloranthus.
Echinocereus neocapillus (Weniger) Blum & Lange in Blum et al. 1998:219.
  = Echinocereus chloranthus var. neocapillus.
Echinocereus neoroemeri* (Mühlenpfordt) Ito 1971:227. nom. illegit. 
  = Echinocereus roemeri (Mühlenpfordt) Rydberg
Wilcoxia nerispina* Backeberg 1960:2078 nom. invalid. 
  = Echinocereus schmollii.
Echinocereus nicholii (Benson) Parfitt 1987:157-158
  = Echinocereus engelmannii var. nicholii.
Echinocereus nicholii fa. guaymasensis* Parker 1996:1 nom. nud.
Echinocereus nicholii ssp. llanuraensis Rutow 1995:61-71.
TL: Mexico, Sonora, Guaymas, 300m (ZSS # A 16936).
Echinocereus nicholii fa. llanuraensis* Parker 1996:1 nom. nud. 
  = Echinocereus nicholii ssp. llanuraensis
Echinocereus nigrispinus* Köhres Catalog 1979. nom. nud.
Echinocereus nivilus* Heyer 1989:26  nom. nud. [= Lau 1101].
Echinocereus nivosus Glass & Foster 1978a:18-19.
TL: Mexico, Coahuila, Southwest of General Cepeda, pass 3 km.
north of El Cinco, Glass & Foster 3764, Feb. 1972 (POM).
Note: Echinocereus albatus* Backeberg 1960:2007 nom. inval.
(no type designated) is usually considered a synonym of this
species.
Echinocereus nivosus fa. albatus* Parker 1996:3 nom. nud.
= Echinocereus albatus*.
Echinocereus noctiflorus* Hort. ex Backeberg 1960:1991 pro syn. 
  = Echinocereus cucumis.
Echinocereus nocturniflorus* Hort. ex Backeberg 1960:1991 pro syn. 
  = Echinocereus cucumis.
Echinocereus nogalensis* Haage 1900:159 nom. nud.
  [Perhaps = Echinocereus pectinatus var. rigidissimus].
Echinocereus ochoterenae Ortega 1928:88-90.
TL: Mexico, Sinaloa, near Concordia, Cerro de la Cobriza, J.G.
Ortega, 24 May 1927 (MEXU?).
Synonyms:
Echinocereus subinermis var. ochoterenae (Ortega) Unger 1984:164.
Echinocereus subinermis ssp. ochoterenae (Ortega) Taylor 1997:10.
Note: Ortega described varieties „A“ and „B“, but did  not give them names.
Echinocereus octacanthus (Mühlenpfordt) Britton & Rose 1922:13.
= Echinopsis octacantha Mühlenpfordt 1848:19.
TL: Northern Texas [San Saba River according to Engelmann],
Roemer.
Synonyms:
Cereus octacanthus (Mühlenpfordt)Coulter 1896:395.
Echinocereus coccineus var. octacanthus* Boissevain & Davidson 1940:39 nom. illegit.
Echinocereus triglochidiatus var. octacanthus* Mühlenpfordt ex Marshall & Bock
1941:118. nom. illegit.
Note: The proper application of this name is uncertain. If it can be shown to be one of
the members of the  Echinocereus coccineus group it would have priority
since it was published first. This would cause considerable nomenclatural changes.
Echinocereus oklahomensis Lahman 1950:128.
[Originally published in Lahman 1935:141 without Latin diagnosis].
TL: Oklahoma, Wichita Mtns., Lahman (MO).
Note: Usually consisdered a synonym of Echinocereus baileyi.
Echinocereus orcuttii Rose ex Orcutt 1926:4.
TL: Mexico, Baja California, Cape Colset, Orcutt (?).
Note: Usually consisdered a synonym of Echinocereus maritimus.
Echinocereus ortegae Rose ex Ortega 1929:24.
TL: Mexico, Sinaloa, Cuenca del Rio Tamazula, Sianori, 800m., Ortega (?).
Note: Taylor (1993) lists Echinocereusscheeri var. koehresianus as a synonym
of this species.
Echinocereus ortegae ssp. koehresianus (Frank) Rischer & Frank 1996:93
    = Echinocereus scheeri var. koehresianus.
Echinocereus pacificus* Haage 1892:118 nom.nud.
Echinocereus pacificus (Engelmann ex Orcutt) Britton & Rose 1922:12.
= Cereus phoeniceus var. pacificus Engelmann ex Orcutt 1886:46.
TL: Mexico, Baja California Norte, Rio San Carlos, Parry et al., 25 Jan. 1883 (MO).
Synonyms:
Cereus pacificus (Engelmann ex Orcutt) Coulter 1896:397.
Echinocereus triglochidiatus var. pacificus (Engelmann ex Orcutt) Engelmann ex 
Marshall & Bock 1941:118.
Echinocereus polyacanthus var. pacificus (Engelmann ex Orcutt) Taylor 1984:160.
Echinocereus polyacanthus ssp. pacificus (Engelmann ex Orcutt) Breckwoldt 1996:15.
Echinocereus pacificus ssp. mombergerianusBlum, Rischer & Rutow in Blum et al. 1998:418. = Echinocereus mombergerianus.
Echinocereus palmeri Britton & Rose 1922:34.
TL: Mexico, Chihuahua, hill near Cd. Chihuahua, E. Palmer 121,
Apr. 1908 (US).
Echinocereus palmeri
ssp. escobedensis Rischer (KuaS 54 (5) 2003)
TL:
Durango, MX. Lau 1565, 18.3.1988 ex cult. coll. Rischer
Echinocereus pamanesiorum Lau 1981:36-41.
TL: Mexico, Zacatecas, Rio Huaynamota, bridge between
Huejuquilla & San Juan Capistrano, Lau 1247 (MEXU).
Echinocereus pamanesiorum ssp. bonatzii (Römer) Römer 1997:45.
= Echinocereus bonatzii.
Echinocereus papillosus Linke ex Rümpler 1885:783.
TL: unknown
Synonyms:
Cereus papillosus (Linke ex Rümpler) Berger 1905:80.
Echinocereus blanckii var. papillosus (Linke ex Rümpler) Benson 1969d:126.
Echinocereus berlandieri var. papillosus (Linke ex Rümpler) Benson 1976:59.
Note: Benson uses the names Echinocereus blanckii var. angusticeps
and Echinocereus berlandieri var. angusticeps for this plant. Usually listed as
Echinocereus papillosus Linke ex Rümpler 1885:783.
Echinocereus papillosus var. angusticeps (Clover) Marshall & Bock 1941:119
= Echinocereus angusticeps.
Echinocereus papillosus ssp. angusticeps* Blum 1997 Seed  List nom. nud.
  = Echinocereus angusticeps.
Echinocereus papillosus var. giganteus* Bailey 1941:270 nom. nud.
Echinocereus papillosus var. rubescens Hildmann ex Schelle
1926:165.
TL: unknown
Echinocereus parkeri Taylor 1988:73.
TL: Mexico, Nuevo Leon, between La Poza & Puerto de los Pastores,
Hansen et al. 3863 (MEXU).
Synonyms:
Echinocereus stramineus var. parkeri* Seed list 1996 –  British Cactus and Succulent 
        Journal.
Echinocereus parkeri ssp. arteagensis Blum & Lange in Blum et al. 1998:175.
TL: Mexico, Coahuila – Nuevo Leon Border, Arteaga Canyon, 2250 m,
        U. Guzman in Sánchez- Mejorada 3807 (MEXU).
Echinocereus parkeri fa. barcena* Parker 1996:3 nom. nud. 
  = Echinocereus barcena.
Echinocereus parkeri var. gonzalezii Taylor 1988:74.
TL: Mexico, Tamaulipas, Sierra de las Vacas, 4km NE of San Jose
d’Llano, 1750m, Gonzalez Medrano 8515 (MEXU).
Echinocereus parkeri ssp. gonzalezii BTaylor) Taylor 1997:9.
   = Echinocereus parkeri var. gonzalezii.
Echinocereus parkeri var. mazapilensis* K. Neumann 1993 nom. prov.
Kaktusblüte p. 55.
Echinocereus parkeri ssp. mazapilensis Blum & Lange in Blum et al. 1998:172.
TL: Mexico, Zacatecas, 0.3 km from El Cobre near Mazapil, 2750 m,
Sánchez – Mejoradaet al. 4041 (MEXU).
Echinocereus paucispinus (Engelmann) Loder 1882:445.
= Cereus paucispinus Engelmann 1856:285.
[fuller description Engelmann 1859:34].
TL: Texas, San Pedro River, Wright, 1849 (MO).
Synonyms:
Echinocereus paucispinus* Engelmann ex Rümpler 1885:794
nom. illegit.
  Echinocereus triglochidiatus var. paucispinus (Engelmann) Engelmann ex
          Marshall & Bock 1941:117.
Echinocereus coccineus var. paucispinus (Engelmann) Ferguson 1989:222.
Echinocereus coccineus ssp. paucispinus Blum, Lange & Rutow
in Blum et al. 1998:433.
           Note:Usually cited as Echinocereus paucispinus Engelmann ex
Rümpler 1885:794
Echinocereus paucispinus var. flavispinus* Haage 1892:118 nom. nud.
Echinocereus paucispinus var. flavispinus* Walton 1899b:18 nom. nud.
Echinocereus paucispinus var. flavispinus Hort. ex Schelle 1926:178.
TL: unknown
Echinocereus paucispinus var. fulvispinus* Haage 1897:12 nom. nud.
Echinocereus paucispinus var. fulvispinus Bowles 1908:28.
TL: unknown
Echinocereus paucispinus var. gonacanthus (Engelmann & Bigelow) Schumann 1897:281
    = Echinocereus gonacanthus.
Echinocereus paucispinus var. hexaedrus (Engelmann & Bigelow) Schumann 1897:281
    = Echinocereus hexaedrus.
Echinocereus paucispinus var. nigrispinus* Haage 1897:12 nom. nud.
Echinocereus paucispinus var. nigrispinus Bowles 1908:28.
TL: unknown
Echinocereus paucispinus var. triglochidiatus* Schumann 1897:281  nom. illegit. 
   = Echinocereus triglochidiatus.
Echinocereus pectinatus (Scheidweiler) Engelmann 1848a:110.
= Echinocactus pectinatus Scheidweiler 1838:492.
TL: Mexico, San Luis Potosi, Villa del Peñasco, Galeotti (?).
Synonyms:
Echinopsis pectinata (Scheidweiler) Fennel 1843:282.
Cereus pectinatus (Scheidweiler) Engelmann 1849:50.
Note:Echinocereus cernosus* Fric 1929:24 nom. nud. is
listed by Backeberg as a synonym of this species.
Echinocactus pectiniferusLemaire 1839:25 is usually
considered a synonym of this species.
Echinocereus pectinatus var. adustus* Haage 1859:19 nom. nud. 
Echinocereus pectinatus var. adustus [mispelled adjustus](Engelmann) Daul 1890:78
= Echinocereus adustus.
Note: Usually ascribed to Schumann 1897:271.
Echinocereus pectinatus fa. adustus (Engelmann) Voss 1894:378 = Echinocereus adustus.
Echinocereus pectinatus var. adustus fa. castaneus (Engelmann) Engelm. ex Schelle
1926:174   = Echinocereus caespitosus var. castaneus.
Echinocereus pectinatus var. armatus* Engelmann ex Seitz 1870:11 nom. nud.
Echinocereus pectinatus var. armatus (Poselger) Poselger ex 
  Rümpler 1885:818.
= Cereus pectinatus var. armatus Poselger 1853:134.
TL: Mexico, Nuevo Leon, near Monterrey. Poselger (?)
Synonyms:
Cereuspectinatus var. spinosus* Coulter 1896:357.
Echinocereus spinosus  Coulter ex Schelle 1926:174 pro syn.
Echinocereus armatus (Poselger) Poselger ex Backeberg & Knuth 1935:318.
Echinocereus caespitosus var. armatus (Poselger) Poselger ex Borg 1937:174.
Echinocereus reichenbachii var. armatus (Poselger) Taylor 1985b:133.
Echinocereus fitchii var. armatus* Mesa Garden Seed Catalog 1996 nom. nud.
Echinocereus reichenbachii ssp. armatus (Poselger) Taylor 1997:9.
Echinocereus fitchii ssp.armatus (Poselger) Blum et al. 1998:314
Echinocereus pectinatus var. bauerii Rother 1923:127.
TL: [no type locality or type mentioned]
Echinocereus pectinatus var. bristolii (Marshall) Marshall1956b:81-82
= Echinocereus bristolii.
Echinocereus pectinatus var. caespitosus (Engelmann) Schumann 1897:272
= Echinocereus caespitosus.
Echinocereus pectinatus var. candicans* Hort. ex Schelle 1907:132 nom. nud.
Echinocereus pectinatus var. candidissimus* Seitz 1878:11 nom. nud.
Echinocereus pectinatus var. castaneus* Seitz 1878:11 nom. nud.
Echinocereus pectinatus var. castaneus (Engelmann) Daul 1890:78
= Echinocereus caespitosus var. castaneus.
Echinocereus pectinatus fa. castaneus (Engelmann) Krainz 1967:50
= Echinocereus caespitosus var. castaneus.
*Echinocereus pectinatus var. centralis Coulter ex Schumann1897:271
error in determination.  = Echinomastus centrilis.
Echinocereus pectinatus var. chloranthus* Haage 1892:118 nom. nud.
Echinocereus pectinatus var. chrysacanthus Schumann 1897:272.
TL: Mexico.
Synonyms:
Echinocereus caespitosus var. chrysacanthus (Schumann)
Schumann ex Borg 1937:174.
Echinocereus pectinatus var. crassispinus Rünge ex Mathsson
1890:465.
TL: Mexico, Coahuila, Saltillo, Mathsson (?).
Echinocereus pectinatus var. ctenoides* Hort. ex Schelle1907:130 pro syn.
    = Echinocereus ctenoides.
Echinocereus pectinatus var. ctenoides* Weniger 1971:31 nom. inval. (no basionym given)
   = Echinocereus ctenoides.
Echinocereus pectinatus var. ctenoides (Engelmann) Weniger ex Frank 1991:135
= Echinocereus ctenoides.
Echinocereus pectinatus ssp. ctenoides (Engelmann) Frank 1997:136
= Echinocereus ctenoides.
Echinocereus pectinatus var. dasyacanthus (Engelmann) Engelmann ex Haage 1892:118
= Echinocereus dasyacanthus.
Echinocereus pectinatus var. dasyacanthus* Earle 1971:80 nom. illegit.  (later homonym)
   = Echinocereus dasyacanthus.
Echinocereus pectinatus var. dasyacanthus* Taylor 1984:179  nom. illegit. (later homonym)
   = Echinocereus dasyacanthus.
Echinocereus pectinatus var. grandicostatus* Schmoll ex Backeberg 1960:2043 nom. nud.
Echinocereus pectinatus var. grandis (Britton & Rose) Marshall 1955:68
= Echinocereus grandis.
Echinocereus pectinatus var. jamauvei *Fric ex Orcutt 1926:4 pro syn. 
  = Echinocereus castaneus.
Echinocereus pectinatus var. labouretianus*  Seitz 1870:11 nom. nud.
Echinocereus pectinatus var. laevior* (Monatsschrift für Kakteenkunde
  index 1912:56) nom. nud.
Echinocereus pectinatus var. minor (Engelmann) Benson 1968:125
for Echinocereus dasyacanthus var. minor*.
    = Echinocereus roetteri.
Echinocereus pectinatus var. neomexicanus (Coulter) Benson 1944:256
= Echinocereus dasyacanthus var. neomexicanus*.
Note: Benson uses this name for Echinocereus dasyacanthus.
Echinocereus pectinatus var. ordustus* Lakerveld Catalog. nom. nud.
Echinocereus pectinatus var. pailanus* Schwarz ex Backeberg 1960:2043 nom. nud.
Echinocereus pectinatus var. pailensis* Bailey 1941:271 nom. nud.
Echinocereus pectinatus var. pictus* Seitz 1870:11 nom. nud.
Echinocereus pectinatus var. rectispinus* Zimmerman1993:277 nom. nud. 
  = Echinocereus dasyacanthus var. rectispinus.
Echinocereus pectinatus var. reichenbachii (Terscheck ex Walpers) Werdermann 1930:t5
= Echinocereus reichenbachii.
Echinocereus pectinatus var. rigidissimus (Engelmann) Engelmann
  ex Rümpler 1885:818.
= Cereus pectinatus var. rigidissimus Engelmann 1856:279.
TL: Arizona, Sierra Pajarito, Schott 1855 (MO).
Synonyms:
Echinocereus pectinatus var. robustior* Hort. ex Hirscht
1897:95 pro syn.
Echinocereus candicans* Runge ex Haage 1900:157 pro syn.
Echinocereus rigidissimus (Engelmann) Hort. ex F. A. Haage 1897, „special
offer“catalog p. 13.
Echinocereus robustior* Britton & Rose 1922:28  pro syn.
Echinocereus dasyacanthus var. rigidissimus (Engelmann) Marshall & Bock 1941:119.
Echinocereus pectinatus fa. rigidissimus (Engelmann) Krainz 1967:50.
Note: Usually listed as Echinocereus rigidissimus Rose 1909a:293.
                Echinocereus nogalensis* Haage 1900:159 nom. nud.  is usually considered a
synonym of this plant.
Echinocereus pectinatus fa. rigidissimus (Engelmann) Krainz 1967:50.
=Echinocereus pectinatus var. rigidissimus.
Echinocereus pectinatus var. robustior* Hort. ex Hirscht 1897:95  pro syn.
  = Echinocereus pectinatus var. rigidissimus.
Echinocereus pectinatus var. robustus Bauer ex Wittmack 1890:513.
TL: Mexico, Sonora, Nogales.
Note: usually considered a synonym of Echinocereuspectinatus var. rigidissimus.
Echinocereus pectinatus var. rotatus (Linke) Linke ex Haage 1892:118
= Echinocereus rotatus.
Echinocereus pectinatus var. rubescens* Weniger 1970:21-22 nom. nud.
Echinocereus pectinatus var. rubispinus Frank & Lau in Frank
1982:32-35.
TL: Mexico, Chihuahua, Cañon de Barbarocas, near Campito in Sierra
Obscura, 1600-1800m, Lau 088 (ZSS AA 18-30).
Synonyms:
Echinocereus rigidissimus var. rubispinus (Frank & Lau) Taylor 1984:175.
Echinocereus rigidissimus ssp. rubispinus Taylor 1997:9.
Echinocereus pectinatus var. rungei* Hort. ex Schelle 1926:174  nom. nud.
Echinocereus pectinatus var. rufispinus* Daul 1890:78 nom. nud.
Echinocereus pectinatus var. rufispinus Schumann 1897:272.
TL: Mexico.
Synonyms:
Echinocereus caespitosus var. rufispinus (Schumann) Schumann ex Borg 1951:221.
Cereus pectinatus fa. rufispinus (Engelmann) Voss 1894:378
= Echinocereus rufispinus.
Echinocereus pectinatus var. scopulorum (Britton & Rose) Marshall 1955:61
= Echinocereus scopulorum.
Echinocereus pectinatus var. sonorensis* (Kaktusy 1973 according toEvanich 1989:111).
Cereus pectinatus var. spinosus* Coulter 1896:357 nom.illegit.
[an unnecessary new name for Echinocereus pectinatus var. armatus].
Echinocereus pectinatus var. steereae* Rümpler ex Weniger 1970:32  pro syn.
   = Echinocereus steereae.
Echinocereus pectinatus var. tamaulipensis Fric ex Schelle 1926:174.
TL: unknown
synonym:
Echinocereus caespitosus var. tamaulipensis (Fric) Fric ex Borg 1937:174.
Echinocereus pectinatus var. texensis* Horner ex Seitz 1870:11 nom. nud.
Echinocereus pectinatus var. texensis Hort. ex Rümpler 1885:817.
TL: Texas
Echinocereus pectinatus var. texensis* Hooker ex Schelle 1907:132  nom. nud.
Echinocereus pectinatus var. wenigeri Benson 1968:124.
TL: Texas, Valverde Co, near Langtry, Benson 16,521 (POM 311,338).
Echinocereus pectinatus ssp. wenigeri (Benson) Blum & Rutow in Blum et
al. 1998: 111=Echinocereus pectinatus var. wenigeri.
Echinocactus pectiniferus Lemaire 1839:25.
TL: unknown
Synonyms:
Cereus pectiniferus (Lemaire) Labouret 1853:320
Note: usually considered a synonym of Echinocereus pectinatus.
Echinocactus pectiniferus var. laevior Monville ex Lemaire 1839:26.
TL: unknown
Synonyms:
Echinopsis pectinata var. laevior (Monville ex Lemaire) Monville ex Förster 1846:365.
*Echinocereus pencilliformis Linke 1858
[of Bolivian origin and therefor not an Echinocereus].
Echinocereus pensilis (K. Brandegge) J. A. Purpus 1908:5.
= Cereus pensilis K. Brandegee 1904:192.
TL: Mexico, Baja California Sur, Sierra de Laguna, T.S. 
    Brandegee 246, Jan/Feb. 1890 (UC).
Synonyms:
Morangaya pensilis (K. Brandegee) Rowley 1974b:44-45.
Echinocereus pentalophus* Engelmann ex Haage 1859:20 nom. nud.
Echinocereus pentalophus* Lemaire 1868:56. nom. nud.
[Probably = Echinocereus pentalophus De Candolle ex Rümpler].
Echinocereus pentalophus (De Candolle) Lemaire ex Rümpler 1885:774.
= Cereus pentalophus De Candolle 1828:117.
TL: Mexico, Hidalgo, near Zimapan, Coulter (?).
Synonyms:
Cereus propinquus* Hort. Genev. ex Walpers 1843:275 pro syn.
Echinocereus pentalophus var. aureovariegatus* Ito 1952:134 nom. nud.
Echinocereus pentalophus var. ehrenbergii (Pfeiffer) Backeberg1960:2003
= Echinocereus ehrenbergii.
Echinocereus pentalophus var. leonensis (Mathsson) Taylor 1985b:78
= Echinocereus leonensis.
Echinocereus pentalophus ssp. leonensis (Mathsson) Taylor 1997:9
= Echinocereus leonensis.
Echinocereus pentalophus
var. leptacanthus* Haage 1859:20 nom. nud.
Echinocereus pentalophus var. leptacanthus* Lemaire ex Rümpler 1885:775 nom. illegit. 
    = Echinocereus leptacanthus.
Echinocereus pentalophus var. procumbens (Engelmann) Fournier 1935:25
= Echinocereus procumbens.
Echinocereus pentalophus fa. procumbens* Parker 1996:4 nom. nud. 
  = Echinocereus procumbens.
Echinocereus pentalophus ssp. procumbens (Engelmann) Blum & Lange
in Blum et al. 1998:129. = Echinocereus procumbens.
Cereus pentalophus var. radicans DeCandolle1828:117.
TL: Mexico, Hidalgo, near Zimapan, Coulter (?).
Synonyms:
Cereus propinquus* Hort. Genev. ex Walpers 1843:275 pro syn.
Echinocereus pentalophus var. radicans* Weniger 1970:51  nom. inval.
Echinocereus pentalophus var. radicans* Weniger 1970:51 nom. inval. 
(no basionym given). = Cereus pentalophus var. radicans.
Cereus pentalophus var. simplex De Candolle 1828:117.
TL: Mexico, Hidalgo, near Zimapan, Coulter (?).
Synonyms:
Cereus propinquus* Salm Dyck ex Otto:1833:366. nom. nud.
Cereus propinquus* De Candolle ex Walpers 1843:333 pro syn.
Echinocereus propinquus De Candolle ex Haage 1892:118.
Echinocereus pentalophus var. simplex* Weniger 1970:51nom. inval.
Echinocereus pentalophus var. simplex* Weniger 1970:51 nom. inval. (no basionym given).
= Cereus pentalophus
var. simplex.
Cereus pentalophus var. subarticulatus De Candolle 1828:117.
TL: Hidalgo, near Zimapan, Coulter (?).
Synonyms:
Cereus propinquus var. subarticulatus* Pfeiffer ex Förster 1846:373 pro syn.
Cereus leptacanthus De Candolle ex Salm-Dyck 1845:27.
Echinocereus leptacanthus Gaillard 1864:209.
Echinocereus pentalophus var. subarticulatus* Weniger 1970:51 nom. inval.
Echinocereus pentalophus var. subarticulatus* Weniger 1970:51 nom. inval.
    (no basionym given).  = Cereus pentalophus var. subarticulatus.
*Echinocereus pentlandii  = Echinopsis pentlandii.
Echinocereus perbellus Britton & Rose 1922:24-25.
TL: Texas, Big Springs, Rose & Standley 12215, 23 Feb.1910 (US 635015, 635016).
Synonyms:
Echinocereus reichenbachii var. perbellus (Britton & Rose) Benson 1969d:127.
Echinocereus caespitosus var. perbellus* Weniger 1970:23 nom. inval.
Echinocereus reichenbachii ssp. perbellus* Blum 1997 Seed List nom. nud.
*Echinocereus persolutus  [from Peru and therefor not an Echinocereus]
Echinocereus phaeacanthus* Cactus & Succulent Journal (U.S.)1951:104 nom. nud.
Echinocereus phoeniceus* Lemaire 1868:57 nom. nud. [Probably = Echinocereus coccineus].
Echinocereus phoeniceus* Engelmann ex Rümpler 1885:788 nom. illegit.
  = Cereus phoeniceus Engelmann 1856:284 which was a new name for  Echinocereus
coccineus
Engelman 1848:94.
Echinocereus phoeniceus* var. albispinus Hort. ex Rümpler 1885:789
Echinocereus phoeniceus* var. brevispinus Engelmann ex Schelle 1907:138 nom. nud.
Cereus phoeniceus ssp. conoideus Engelmann 1857:37
  = Echinocereus conoideus.
Echinocereus phoeniceus* var. conoideus Schumann 1896:150
= Echinocereus conoideus.
Echinocereus phoeniceus* var. densus Schelle 1907:138 nom. nud.
Echinocereus phoeniceus* var. inermis Schumann 1896:150
  (see Echinocereus coccineus var. inermis).
Echinocereus phoeniceus* var. longispinus Hort. ex Rümpler 1885:789
Cereus phoeniceus var. pacificus Engelmann ex Orcutt 1886:46
(see Echinocereus pacificus).
Echinocereus phoeniceus* var. rufispinus Hort. ex Rümpler 1885:789
Echinocereus phoeniceus* var. utahensis* Schelle 1907:138 nom. nud.
Echinocereus pleiogonus Croucher 1878:290.
TL: unknown [perhaps an Echinopsis]
Echinocereus pleiogonus* Labouret ex Rümpler 1885:828-829  nom. illegit. (later homonym).
= Cereus pleiogonus Labouret 1853:317.
TL: unknown
Echinocereus plomosus* Köhres Catalog 1984. nom. nud. 
[= Lau 768 from Plomosos, Coahuila ]
Echinocereus polichephalus* Daul 1890:78 nom. nud.
Echinocereus polycephalus
* Hort. Berol. ex Haage 1897 nom. nud.
Echinocereus polyacanthus Engelmann 1848a:104.
TL: Mexico, Chihuahua, Cosihuiriachi, Wislizenus, 1846 (MO).
Synonyms:
Cereus polyacanthus (Engelmann)Engelmann 1849:50.
Echinocereus triglochidiatus var. polyacanthus (Engelmann) Benson 1944:253.
Echinocereus polyacanthus var. acifer* Parker 1996:4 nom. nud. 
  = Echinocereus acifer.
Echinocereus polyacanthus ssp. acifer (Otto ex Salm-Dyck) Taylor 1997:9.
  = Echinocereus acifer.
Echinocereus polyacanthus var. acifer fa. huitcholensis* Parker 1996:4. nom. nud.
= Echinocereus huitcholensis.

Echinocereus polyacanthus var. acifer fa. matthesianus* Parker 1996:4 nom. nud.
= Echinocereus matthesianus*

Echinocereus polyacanthus var. albispinus Hort. ex Schelle 1926:179.
TL: unknown
Echinocereus polyacanthus var. bergeanus Fobe 1911:56.
TL: unknown
Echinocereus polyacanthus var. bergeanus* Hort ex Schelle  1926:179 nom. illegit.
(later homonym).
Echinocereus polyacanthus var. densus (Regel) Taylor 1984:159
= Echinopsis valida var. densa Regel 1852:295.
Note: This name is used by Taylor (1984) as the name of Echinocereus  acifer.
Echinocereus polyacanthus var. durangensis* Parker 1996:4 nom. nud.
= Echinocereus durangensis.
Echinocereus polyacanthus var. durangensis fa. marksianus* Parker 1996:4 nom. nud.
=Echinocereus marksianus.
Echinocereus polyacanthus var. galtieri (Rebut ex Fobe) Schelle 1926:179
= Echinocereus galtieri.
Echinocereus polyacanthus var. huitcholensis (Weber) Taylor 1988:82
= Echinocereus huitcholensis.
Echinocereus polyacanthus ssp. huitcholensis (Weber) Taylor 1997:9
=  Echinocereus huitcholensis.
Echinocereus polyacanthus var. leeanus* Parker 1996:4 nom. nud. 
  = Echinocereus leanus.
Echinocereus polyacanthus var. longispinusHort. ex Schelle 1926:179.
TL: unknown.
Echinocereus polyacanthus var. mombergerianus* Parker 1996:4 nom. nud. 
  = Echinocereus mombergerianus.
Echinocereus polyacanthus ssp. mombergerianus (Frank)Breckwoldt & Matylewicz
1996:15      = Echinocereus mombergerianus.
Echinocereus polyacanthus var. neomexicanus* Weniger 1970:44 nom. inval.
    (no basionym given)     = Echinocereus neomexicanus.
Echinocereus polyacanthus var. nigrispinus Fobe 1911:55.
TL: unknown
Echinocereus polyacanthus var. nigrispinus* Hort. ex Schelle
1926:179 nom. illegit. (later homonym).
Echinocereus polyacanthus var. pacificus (Engelmann ex Orcutt)
Taylor 1984:160     = Echinocereus pacificus.
Echinocereus polyacanthus ssp. pacificus (Engelmann ex Orcutt) Breckwoldt
in Breckwoldt & Matylewicz 1996:15
    = Echinocereus pacificus.
Echinocereus polyacanthus var. rosei* Weniger 1970:43 nom. inval. 
  (no basionym given)     = Echinocereus rosei.
Echinocereus polyacanthus var. rufispinus Fobe 1911:55-56.
TL: unknown
Echinocereus polyacanthus var. rufispinus* Hort. ex Schelle 1926:179 nom. illegit.
   (later homonym).
Echinocereus polycephalus* Blanc 1888:39 nom. nud.
Echinocereus polycephalus* Hort. Berol. ex Haage 1900:160 nom. nud.
Echinocereus polycephalus* var. albispinus* Fric 1924:121 nom. nud.
Echinocereus polycephalus* var. brevinigrispinus* Fric 1924:121
nom. nud.
Echinocereus polycephalus* var. nigrispinus* Fric 1924:121 nom. nud.
Echinocereus polycephalus* var. spinosissimus* Fric 1924:121 nom. nud.
Echinocereus polycephalus var. xphoeniceus* Fric 1924:121 nom. nud.
Echinocereus polypus* Fric 1924:121 nom. nud.
Echinocereus poselgeri Lemaire 1868:57
[A new name needed because of earlier homonym].
= Cereus tuberosus Poselger 1853:131 nom. illegit. 
(later homonym) [not Cereus tuberosus Pfeiffer 1837.]
TL: Texas, near Rio Grande River, above Belleville, Poselger, 
1850 (MO).
Synonyms:
Echinocereus tuberosus* Rümpler 1885: 131 nom. illegit.
          Cereus poselgeri Coulter 1896:398.
Wilcoxia poselgeri (Lemaire) Britton & Rose 1909:434.
Wilcoxia tuberosa*  Berger 1929:134 nom. illegit.
Echinocereus poselgeri fa. australis* Parker 1996:6 nom. nud.
Echinocereus poselgeri ssp. kroenleinii (Cartier) Lange 1995:138.
= Wilcoxia kroenleinii Cartier 1980 (Succulentes 2(2):2-3).
[see Echinocereus kroenleinii].
Echinocereus poselgeri fa. kroenleinii* Parker 1996:6 nom. nud. 
  = Echinocereus kroenleinii.
Echinocereus poselgeri fa. tamaulipensis* Parker 1996:6 nom. nud.
= Echinocereus tamaulipensis.
Echinocereus poselgeri fa. tuberosa* Parker 1996:6 nom. nud.
Echinocereus poselgeri var. waldeisii* Parker 1996:6 nom. nud.
= Echinocereus waldeisii. 
Echinocereus poselgeri var. waldeisii fa. deherdtii* Parker 1996:6 nom. nud. 
   = Echinocereus tamaulipesis var. deherdtii.
Echinocereus poselgerianus Linke 1857:239.
TL: Mexico
Synonyms:
Cereus poselgerianus (Linke) Berger 1905:80.
Echinocereus blanckii var. poselgerianus (Linke) Backeberg 1960:1999.
Note: usually considered a synonym of Echinocereus berlandieri.
Echinocereus primolanatus* Schwarz ex Backeberg 1960:2043 nom.
   inval.  (no type designated).
Echinocereus primolanatus Taylor 1985b:130.
TL: Mexico, Coahuila, Sierra Paila, north of Estacion Marte,
cultivated by D. Parker, 1985(K).
Synonyms:
Echinocereus primolanatus* Schwarz ex Backeberg 1960:2043
        nom. inval. (no type designated).
Echinocereus princeps Föster 1861:163.
TL: unknown [Of unknown origin and perhaps from South America in
which case it would not be an Echinocereus].
Echinocereus procumbens* Lemaire 1868:56 nom. nud.
[Probably = Echinocereus procumbensEngelmann ex Rümpler].
Echinocereus procumbens (Engelmann) Engelmann ex Rümpler 1885:781.
= Cereus procumbens Engelmann 1849:50.
TL: Mexico, Tamaulipas, Burita at mouth of Rio Grande, St. Louis
Volunteers 1846 (MO).
Synonyms:
Echinocereus pentalophus var. procumbens (Engelmann) Fournier 1935:25.
Echinocereus pentalophus fa. procumbens* Parker 1996:4 nom. nud.
        Echinocereus pentalophus ssp. procumbens (Engelmann) Blum & Langein Blum et al.
1998:129. 

Echinocereus procumbens var. gracilior Dautw. ex Schelle 1926:165
TL: unknown
Echinocereus procumbens var. longispinus Hort. ex Hirscht 1902:135.
TL: unknown
Cereus propinquus Salm Dyck ex Otto 1833:366.
  = Echinocereus pentalophus var. simplex.
Echinocereus propinquus De Candolle ex Haage 1892:118
  = Echinocereus pentalophus var. simplex.
Cereus propinquus var. subarticulatus* Pfeiffer ex Förster 1846:373 pro syn.
    = Echinocereus pentalophus var. subarticulatus.
Echinocereus pseudohempelii* Künzler Cat. nom. nud. 
  = Echinocereus kuenzleri.
Echinocereus pseudopectinatus (Taylor) Taylor 1989:74.
= Echinocereus bristolii var. pseudopectinatus Taylor 1985b:120.
TL: Mexico, Sonora, Moctezuma, Lau 607, 1973, cultivated by Taylor, 21 June 1984 (K).
Synonyms:
Echinocereus scopulorum ssp. pseudopectinatus* Blum & Langein Blum et al. 1998:342.
Echinocereus pubellus* Cactus and Succulents Journal (US) 7:5 1936 nom. nud
Echinopsis pulchella var.flore kermesina* Haage ex Förster1846:364. pro syn. 
    = Echinocereus amoenus.
Echinopsis pulchella var. rosea* Labouret 1853:292. pro syn. 
  = Echinocereus amoenus.
Echinocereus pulchellus (Martius) F. Seitz 1870:11.
[see Taylor 1989:75 ].
= Echinocactus pulchellus Martius 1832:342.
TL: Mexico, Hidalgo, Pachuca, Karwinsky (?).
Synonyms:
Cereus pulchellus Pfeiffer 1837:74.
Echinonyctanthus pulchellus (Martius) Lemaire 1839:85.
Echinopsis pulchella (Martius) Zuccarini ex Förster 1846:363.
Note: Usually listed as Echinocereus pulchellus Schumann 1894:185.
Echinocereus pulchellus var. acanthosetus Arias & Guzman in Arias Montes
et al. 1997:63.
TL: Mexico, Oaxaca, between Magdalena Jicotlan & Tepelmeme de
Morelos, Arias, Guzman-Cruz & Gama 958, 29 Oct. 1991 (MEXU).
Synonyms:
Echinocereus pulchellus ssp. acanthosetus (Arias & Guzman)Blum in Blum et al.
         1998:261.
Echinocereus pulchellus ssp. acanthosetus (Arias & Guzman) Blum in
Blum et al. 1998:261. = Echinocereus pulchellus var. acanthosetus
Echinocereus pulchellus var. amoenus (Dietrich) Förster ex Schumann 1897:253
= Echinocereus amoena.
Echinocereus pulchellus ssp. amoenus* Blum 1997 Seed List nom. nud. 
  = Echinocereus. amoenus.
Echinocereus pulchellus var. costatus Halda , Kupčák & Sladk.2000 in Acta Mus. Richnov.
Sect. Nat., 7(1): 34 (2000)
TL: San Luis Potosí/Zacatecas border;  J.J. Halda; J. Sladkovský 8504131 (PR)
Echinocereus pulchellus var. sharpii Taylor 1989:75-77.
TL: Mexico, Nuevo Leon, near San Roberto, 1900m, 24º50´N 100 25´W,
Gonzalez G. 1985 (seed), cult. N. P. Taylor, 4 May 1988 (K).
synonyms:
Echinocereus pulchellus ssp. sharpii (Taylor) Taylor 1997:9
Echinocereus sharpii* S & K Breckwoldt 1998:1. nom. nud.
Echinocereus pulchellus ssp. sharpii (Taylor) Taylor 1997:9
 = Echinocereus pulchellus var. sharpii.
Echinocereus pulchellus subsp. sladkovskyi Halda & Kupčák 2000.
Acta Mus. Richnov. Sect. Nat., 7(1): 34 (2000)
    TL: Hidalgo, MX ; J.J. Halda; J. Sladkovský 850418033 (PR)
Echinocereus pulchellus ssp. venustus Blum & Rischer in Blum et al. 1998:255.
    TL: Mexico San Luis Potosi [Lau 1036] W. Rischer, 5 Dec. 1974 (ZSS A 17984).
Echinocereus pulchellus var. weinbergii (Weingart) Taylor 1985a:272
= Echinocereus weinbergii.
Echinocereus pulchellus ssp. weinbergii (Weingart) Taylor 1997:9
  = Echinocereus weinbergii.
Echinocereus purpureus Lahman 1935:141-143.
TL: Oklahoma, near Medicine Park, Lahman (MO).
Synonyms:
Echinocereus caespitosus var. purpureus* Weniger 1970:23  nom. inval.
Echinocereus reichenbachii var. purpureus* Parker 1996:5 nom. nud.
Echinocereus radians Engelmann 1848a:104.
TL: Mexico, Chihuahua, Cosihuiriachi, Wislizenus, 1846 (MO).
Synonyms:
Cereus adustus var. radians (Engelmann) Engelmann 1849:50.
Echinocereus adustus var. schwarzii fa. radians* Parker 1996:6 nom. nud.
Note: Usually considered a synonym of Echinocereus adustus.
Echinocereus raphicephalus Förster 1861:164.
TL: unknown. [May be of South American origin and not an Echinocereus]
Echinocereus rayonesensis Taylor 1988:75-76.
TL: Mexico, Nuevo Leon, Rayones, Lau 1101 (K).
Echinocereus rayonesensis fa. barkeanus* Parker 1996:3 nom. nud.
[Probably = Echinocereus barkeanus*]
Echinocereus rectispinus Peebles 1938:675.
TL: Arizona, hills near Nogales, 3900 ft., Peebles SF 905
(US 1729266).
Synonyms:
Echinocereus fendleri var. rectispinus (Peebles) Benson 1944:259.
Echinocereus fasciculatus var. rectispinus* Benson 1969d:131 pro syn.
Echinocereus fendleri ssp. rectispinus (Peebles) Taylor 1997:9
Echinocereus rectispinus var. robustus Peebles 1938:675
(see Echinocereus robustus Peebles).
Echinocereus reichenbachianus* Linke 1857:83 nom. nud.
Echinocereus reichenbachianus Hort. ex Haage 1859:20.
[See Echinocereus reichenbachii].
= Echinocactus reichenbachianus (Terscheck) Terscheck ex Fennel
1843:282 (Terscheck, Verzeichnis 3. According to Schumann
1897:271).
Synonyms:
Echinopsis pectinata var. reichenbachiana (Terscheck) Salm-Dyck 1845:26.
Cereus reichenbachianus (Terscheck) Labouret 1853:318.
Echinocereus reichenbachianus* Engelmann ex Haage 1893:23
Note: See note under Echinocereus reichenbachii.
Echinocereus reichenbachianus* Engelmann ex Seitz 1878:11 nom. nud. 
Echinocereus reichenbachianus* Engelmann ex Haage 1893:23
(according to Index Kewensis 2:813) nom. illegit. (later homonym).
Cereus reichenbachianus var. castaneus (Engelmann) Labouret
1853:319 = Echinocereus caespitosus var. castaneus.

Las Cruces, New Mexico 1 May 2006. Permission to copy or reprint
for noncommercial purposes is granted.

Echinocereus reichenbachii – Echinocereus zuniensis

Echinocereus reichenbachii (Terscheck ex Walpers) Haage Jr. ex Britton & Rose 1922:25.
= Echinocactus reichenbachii Terscheck ex Walpers 1843:320
(Terscheck Verzeichnis 2. According to Schumann 1897:452).
TL: Mexico.
Synonyms:
Echinocereus pectinatus var. reichenbachii (Terscheck ex Walpers) Werdermann 1930:t5.
Echinocereus caespitosus var. reichenbachii* Ters. ex Borg 1937:174 nom. illegit.
Note: Since Terscheck’s Original publication has not been found there is confusion
concerning whether the two names Echinocactus reichenbachii and Echinocactus 
               reichenbachianus refer to the same plant. The name  Echinocereus reichenbachii is usually
applied to Echinocereus caespitosus although this is uncertain since the original description
of Echinocactus reichenbachii did not have a description of the plant’s flower, and no
collector or exact type locality is listed. Benson (1982:946) designated Palmer 511, near
Saltillo, Mexico, 10 April 1905 (US 570013) as the neotype. Unless someone is able to
locate a copy of Terscheck’s original publication questions will remain concerning the
proper use of this name.
Echinocereus reichenbachii var. albertii Benson 1969d:127.
TL: Texas, Jim Wells Co., near Alice, Benson 16,550 (POM
317080).
Synonyms:
Echinocereus reichenbachii var. fitchii fa. albertii* Parker 1996:5 nom. nud.
Echinocereus fitchii var. albertii* Mesa Garden Seed Catalog1996 nom. nud.
      Echinocereus fitchii ssp. alberti (Benson) Blum & Lange in Blum et al. 1998:313.
Note: Echinocereusmelanocentrus* Lowry 1936:20 is usually considered a synonym
of this variety.
Echinocereus reichenbachii var. albiflorus* Seela 1940:92 nom. nud.
Echinocereus reichenbachii var. albispinus (Lahman) Benson1969d:127
= Echinocereus albispinus Lahman 1935:143.
Note: Benson uses this name instead of Echinocereus baileyi.
Echinocereus reichenbachii var. anigosanthus* Ito 1981. nom. nud.
Echinocereus reichenbachii var. armatus (Poselger) Taylor 1985b:133
= Echinocereus pectinatus var. armatus.
Echinocereus reichenbachii ssp. armatus (Poselger) Taylor 1997:9
Echinocereus pectinatus var. armatus.
Echinocereus reichenbachii var. aureiflorus* Seela 1940:92 nom. nud.
Echinocereus reichenbachii var. baileyi (Rose) Taylor 1985b:133
= Echinocereus baileyi.
Echinocereus reichenbachii ssp. baileyi (Rose) Taylor 1997:9
  = Echinocereus baileyi.
Echinocereus reichenbachii var. baileyi fa. albispinus* Mesa Garden Seed Catalog 1996 nom. nud. 
    = Echinocereus albuspinus.
Echinocereus reichenbachii  subsp. burrensis G.Frank , Metorn & E.Scherer 2000,
Echinocereenfreund 13(3): 77 (63-77; figs.).
TL: Serranias del Burro, Coahuila, MX; G. R. W. Frank 042 (ZSS)
Echinocereus reichenbachii var. caespitosus* Mesa Garden Seed Catalog 1996 nom. nud.
Echinocereus reichenbachii fa. caespitosus* Parker 1996:5 nom. nud. 
  = Echinocereus caespitosus.
Echinocereus reichenbachii ssp. caespitosus* Blum & Lange in Blum et al.1998:324.
  = Echinocereus caespitosus.
Echinocereus reichenbachii var. chisoensis (Marshall) Benson 1969d:127
= Echinocereus chisoensis.
Echinocereus reichenbachii var. fitchii (Britton & Rose) Benson 1969d:127
= Echinocereus fitchii.
Echinocereus reichenbachii ssp. fitchii (Britton & Rose) Taylor 1997:9.
= Echinocereus fitchii.
Echinocereus reichenbachii var. fitchii fa. albertii* Parker 1996:5 nom. nud.
= Echinocereus reichenbachii
var. albertii
Echinocereus reichenbachii var. fitchii fa. melanocentrus* Parker 1996:5 nom. nud.
= Echinocereus melanocentrus*

Echinocereus reichenbachii fa. pailanus* Parker 1996:5 nom. nud.
Echinocereus reichenbachii var. perbellus (Britton & Rose) Benson
1969d:127 = Echinocereus perbellus.
Echinocereus reichenbachii ssp. perbellus* Blum 1997 Seed List nom. nud.
= Echinocereus perbellus.

Echinocereus reichenbachii var. purpureus* Parker 1996:5 nom. nud. 
  = Echinocereus purpureus.
*Echinocereus reichii (from Chile and therefor not an Echinocereus).
Echinocereus rigidispinus* Engelmann ex Haage 1859:20 nom. nud.
Echinocereus rigidissimus Hort. ex F. A. Haage 1897:13.
  = Echinocereus pectinatus var. rigidissimus.
Echinocereus rigidissimus* Rose 1909a:293 nom. illegit.(later homonym)
    = Echinocereus pectinatus var. rigidissimus.
Echinocereus rigidissimus var. rubispinus (Frank & Lau) Taylor 1984:175
    = Echinocereus pectinatus var. rubispinus Frank & Lau 1982:32-35.
Echinocereus rigidissimus ssp. rubispinus (Frank & Lau) Taylor 1997:9.
    = Echinocereus pectinatus var. rubrispinus.
Echinocereus ritteri* Haage 1958:182 nom. nud.[A form of Echinocereus pentalophus]
Echinocereus robustior* Britton & Rose 1922:28 pro syn.
  = Echinocereus pectinatus var. rigidissimus.
Echinocereus robustus (Peebles) Peebles 1940:219.
= Echinocereus rectispinus var. robustus Peebles 1938:675.
TL: Arizona, Pima Co. Between Tucson & Sabino Canyon, Peebles SF896, 27 Aug 1935
(US 1729267).
Synonyms:
Echinocereus fendleri var. robustus* Benson 1944:259 nom. illegit. (later homonym).
Echinocereus fasciculatusfa. robustus* Parker 1996:2  nom. nud. 
Note: This plant is usually treated as a synonym of Echinocereus fasciculatus.
Echinocereus roemeri* Lemaire 1868:56 nom. nud.
Echinocereus roemeri (Engelmann) Engelmann ex Haage 1859:20.
= Cereus roemeri Engelmann 1849:50. [not Mühlenpfordt].
TL: Texas, granitic region of the Llano. Lindheimer, 1847 (MO).
Synonyms:
Echinocereus coccineus fa. roemeri* Parker 1996:4 nom. nud.
Echinocereus coccineus ssp. roemeri Blum, Lange & Rutow in Blum et al.1998:433.
Note: Usually listed as Echinocereus roemeri Engelmann ex Rümpler 1885:792.
Echinocereus roemeri*(Mühlenpfordt) Rydberg 1906:146 nom. illeg. 
  (later homonym).
= Cereus roemeri Mühlenpfordt 1848:19. [not Engelmann].
TL: Northern Texas, Roemer (?).
Echinocereus roetteri (Engelmann) Engelmann ex Rümpler 1885:829-830.
= Cereus roetteri Engelmann 1856:345. [fuller description
Engelmann 1859:33, t. 41, fig. 3-5].

TL: Texas, stony hills near Frontera [El Paso], Wright, 1851 (MO).
Synonyms:
Cereus dasyacanthus var. minor* Engelmann 1856:278.
Echinocereus pectinatus var. minor (Engelmann) Benson 1968:125.
Echinocereus dasyacanthus fa. roetteri* Parker 1996:2 nom. nud.
Note: This plant is probably a natural hybrid.
Echinocereus roetteri var. lloydii (Britton & Rose) Backeberg 1960:2027
= Echinocereus lloydii.
Echinocereus roetteri var. neomexicanus ( J.M.Coult. ) A.D.Zimmerman,
Cacti Trans-Pecos 211. 2004 pro sp. hybr.
= Echinocereus pectinatus var. neomexicanus (J.M.Coult.) L.D.Benson 1944
Echinocereus roseanus* Backeberg 1960:2071 pro syn. 
  = Echinocereus rosei.
Echinocereus rosei Wooten & Standley 1915:457.
TL: New Mexico, Doña Ana Co., hills near Agricultural College
[Las Cruces], Standley 1235, 1907 (US 535093).
Synonyms:
Echinocereus triglochidiatus var. rosei (Wooten & Standley) Marshall 1950:63.
Echinocereus roseanus* Backeberg 1960:2071 pro syn.
Echinocereus polyacanthus var. rosei* Weniger 1970:44  nom. inval.
Echinocereus coccineus fa. rosei* Blum 1990:13  nom. inval.
Echinocereus triglochidiatus var. neomexicanus fa. rosei* Blum  1990:12. pro syn.
Echinocereus coccineus var. neomexicanus fa. rosei* Parker 1996:4 nom. nud.
Echinocereus coccineus ssp. rosei Blum & Rutow in Blum  et al. 1998:426.
Note: This plant is often listed under the name Echinocereus triglochidiatus var. neomexicanus
            the type of which is probably a natural hybrid.
Echinocereus rosei var. jarillensis* Künzler Cat. nom. nud.
Echinocereus rotatus Linke 1858:85.
TL: South Texas.
Synonyms:
Echinocereus pectinatus var. rotatus (Linke) Linke ex Haage 1892:118.
Note: This name is usually consisdered a synonym of Echinocereus caespitosus.
Echinocereus rubescens Dams 1905:92 -93.
TL: unknown
Note: This species is often considered to be a synonym of Echinocereus dasyacanthus.
Echinocereus rufispinus Engelmann 1848a:104-105.
TL: Mexico, Chihuahua, Cosihuiriachi, Wislizenus, 1846 (MO).
Synonyms:
Cereus rufispinus (Engelmann)Engelmann 1849:50.
Cereus pectinatus fa. rufispinus (Engelmann) Voss 1894:378.
Echinocereus adustus var. schwarzii fa. rufispina* Parker 1996:6 nom. nud.
Note: Usually considered a synonym of Echinocereus adustus.
Echinocereus rungei Schumann 1895:124.
[This is a new name for Echinocereus texensis* Runge]
= Echinocereus texensis* Runge 1894:61. nom. illegit.(later homonym).
TL: Texas, Runge. (?).
Note: The plant is usually considered a synonym of Echinocereus papillosus.
Echinocereus runyonii Orcutt 1926:5.
TL: Texas, mouth of Rio Grande River, Orcutt 1048, 1924 (?).
Note: This plant is probably the same as Echinocereus  procumbens.
Echinocereus russanthus Weniger 1969:41-42.
TL: Texas, Brewster Co., Christmas Mtns., Weniger 712, (UNM).
Nomencatural synonyms:
Echinocereus chloranthus var. russanthus (Weniger) Lamb ex Rowley 1974:7.
Echinocereus viridiflorus var. russanthus (Weniger) Leuck 1980:34.
Note: Echinocereus chloranthus var. vulpis-cauda* is probably a synonym of this species
Echinocereus russanthus var. cowperi* Hort. [HK 1287, from Cooks Range, Luna County,
New Mexico ]. = Echinocereus chloranthus  ssp. rhyolithensis 
Echinocereus russanthus var. fiehnii Trocha 1997:65-69.
TL: Mexico, Chihuahua, Cañon de Santa Clara, Sierra del Nido, Lau 1076
      (Universität Hamburg No. 1066)
Synonyms:
Echinocereus russanthus var. finnii* Köhres Catalog 1981. nom. nud.
Echinocereus finnii* Hort. ex Taylor 1985 101 pro syn.
Echinocereus fiehnii* Lau 1992:74
Echinocereus russanthus fa. finnii* Parker 1996:3 nom. nud. 
      Echinocereus russanthus ssp. fiehnii(Trocha)Blum & Lange in  Blum et al.1998:212.
Echinocereus russanthus ssp. fiehnii(Trocha)Blum & Lange in Blum et al.1998:212.
  = Echinocereus russanthus var. fiehnii.
Echinocereus russanthus var. finnii* Köhres Catalog 1981. nom. nud.
= Echinocereus russanthus var. fiehnii.
Echinocereus russanthus fa. finnii* Parker 1996:3 nom. nud.
= Echinocereus russanthus var. fiehnii.
Echinocereus russanthus ssp. milleri* Blum 1997 Seed List nom. nud. 
  = Echinocereus milleri
Echinocereus russanthus var. neocapillus* Mesa Garden Seed Catalog 1996 nom. nud.
    = Echinocereus chloranthus var. neocapillus.
Echinocereus russanthus var. vulpis-cauda* Köhres Catalog 1981 nom. nud.
= Echinocereus chloranthus var. vulpis-cauda*
Echinocereus russanthus fa. vulpis-cauda* Parker 1996:3 nom. nud. 
  = Echinocereus chloranthus var. vulpis-cauda*
Echinocereus russanthus var. weedinii* Mesa Garden Seed Catalog 1996 nom. nud. 
    = Echinocereus viridiflorus var. weedinii.
Echinocereus russanthus ssp. weedinii Leuck ex Blum & Lange in Blum et al. 1998:214-216.
    = Echinocereus viridiflorus var. weedinii.
Echinocereus salm-dyckianus Scheer 1856:291.
TL: Mexico, Chihuahua.
Synonyms:
Cereus salm-dyckianus (Scheer) Hemsley 1880:545.
Echinocereus scheeri var. salm-dyckianus* Anett ex Parker 1996:4 nom. nud.
Echinocereus salm-dyckianus ssp. bacanorensis Rischer & Trocha 1998:3-10.
TL: Mexico, Sonora, near Bacanora, Trocha AX18239 (ZSS).
Echinocereus salm-dyckianus var. gracilior* Haage 1900:161 nom. nud.
Echinocereus salm-dyckianus var. gracilior Hort. ex Schelle 1926:163.
TL: unknown
Echinocereus salm-dyckianus var. noctiflorusHeid 1944:27.
TL: Mexico, Chihuahua, Cerocauhui, R.T. Craig (CA 305550).
Echinocereus salm-dyckianus ssp. obscuriensis Blum 1994:79-83.
= Echinocereus scheeri var. obscuriensis.
Echinocereus salmianus Hort. ex Rümpler 1885:809.
TL: unknown
Synonyms:
Cereus salmianus (Hort. ex Rümpler) Weber in Bois 1894:279.
Echinocereus scheeri fa. salmianus* Parker 1996:4 nom. nud.
Echinocereus saltillensis* Mats. ex Haage 1900:161 nom. nud.
Echinocereus sanborgianus (Coulter) Schumann 1897:174.
= Cereus sanborgianus Coulter 1896:391.
TL: Mexico, Baja California, rocky table lands south of San
Borja, Gabb 9, 1867 (MO).
Synonyms:
Echinocereus brandegeei fa. sanborgianus* Parker 1996:1 nom. nud.
Note: Usually considered a synonym of Echinocereus brandegeei.
Echinocereus sandersii Orcutt 1926:5.
TL: California, Black Canyon, Providence Mtns.5000ft, Gibson & Sanders 1925 (?).
Note: Usually considered a synonym of Echinocereus mojavensis.
Echinocereus sangre de christo Schumann 1897:278.
TL: Mexico, Mexico, near Pachuca.
Echinocereus sanguineus Blanc 1890:58.
TL: unknown
Note: Probably a form of Echinocereus viridiflorus.
Echinocereus sanpedroensisRaudonat & Rischer 1995:88-97.
TL: Mexico, Sonora, Rancho San Pedro, Rischer, May 1995 (ZSS).
Note: Echinocereus subterraneus* Backeberg 1960:2012 is sometimes
listed as a synonym of this plant.
Echinocereus santaritensisBlum & Rutow in Blum et al.
1998:373-375.
TL: Airzona, Santa Rita Mountains, Rutow 52-56, 16 Apr. 1996 (UA).
Echinocereus sarissophorus Britton & Rose 1922:38-39.
TL: Mexico, Coahuila, near Saltillo, Palmer 100, Apr. 1898 (US).
Synonyms:
Echinocereus enneacanthus fa. sarissophorus* Parker 1996:2
nom. nud.
Echinocereus scheeri* Lemaire 1868:57 nom. nud.
      [Probably = Echinocereus scheeri Salm-Dyck ex Scheer].
Echinocereus scheeri (Salm-Dyck) Salm-Dyck ex Scheer 1856:291
= Cereus scheeri Salm-Dyck 1850:190-191.
TL: Mexico, Chihuahua, Potts (?).
Synonyms:
Echinocereus schlini* Britton & Rose 1922:43 pro syn.
Echinocereus scheeri var. australis* Taylor 1988:80 pro syn.
  = Echinocereus scheeri var. koehresianus.
Echinocereus scheeri var. compactus* Hort. ex Haage 1897:13 nom. nud.
Echinocereus scheeri var. gentryi (Clover) Taylor 1984:154
= Echinocereus gentryi.
Echinocereus scheeri ssp. gentryi (Clover) Taylor 1997:9
  = Echinocereus gentryi.
Echinocereus scheeri var. gentryi cultivar ‚Cucumis‘ (Clover) Taylor 1984:155.
= Echinocereus cucumis.
Echinocereus scheeri var. gentryi fa. cucumis* Parker 1996:4  nom. nud.  = Ecinocereus cucumis.
Echinocereus scheeri var. gracilior Hildmann ex Schelle 1926:162.
TL: unknown
Echinocereus scheeri var. koehresianus Frank 1988:186-189.
TL: Mexico, Durango, between El Carrizo & Loberas, Lau 1143 (ZSS).
Synonyms:
Echinocereus koehresianus* Hort. ex Taylor 1988:80 pro syn.
Echinocereus koehresii* Hort. ex Taylor 1988:80 pro syn.
Echinocereus scheeri var. australis* Taylor 1988:80 pro syn.
Echinocereus ortegae ssp. koehresianus (Frank) Rischer & Frank 1996:93.
Echinocereus ortegae fa. koehresianus* Parker 1996:4 nom. nud.
Note: Taylor (1993) lists this variety as a synonym of Echinocereus ortegae.
Echinocereus scheeri var. major Rother 1905:175.
TL: unknown
Echinocereus scheeri var. minor Rother 1905:175.
TL: unknown
Echinocereus scheeri var. nigrispinus* Scheer 1856:291 nom. nud.
Echinocereus scheeri var. obscuriensis Lau 1989:34-36.
TL: Mexico, Chihuahua, Sierra Obscura, Lau 091 (ZSS).
Synonyms:
Echinocereus salm-dyckianus ssp. obscuriensis(Lau) Blum1994:79.
Echinocereus scheeri var. salm-dyckianus fa. obscuriensis* Parker 1996:4 nom. nud.
Echinocereus scheeri ssp. paridensis Rischer & Trocha 1998:58-65
TL:Mexico , Sinaloa, Near Parida,  Rischer 1992/312 (ZSS AX 18719)
Echinocereus scheeri ssp. polyacanthoides*  Momberger 1994:58.
nom. prov. (from Tomochic, Chihuahua, Mexico)
Echinocereus scheeri ( Salm-Dyck ) Scheer subsp. rischeri R.C.Römer 2004,
Echinocereenfreund 17(1): 8 (3-13; figs.). 2004
TL: Municipio Guachochi, Cumbres de Sinforosa, MX;  Werner Rischer WR 605 (UAT).
Echinocereus scheeri var. robustior Hort. ex Schelle 1926:162.
TL: unknown
Echinocereus scheeri var. salm-dyckianus* Anett ex Parker 1996:4  nom. nud.
= Echinocereus salm-dyckianus.
Echinocereus scheeri var. salm-dyckianus fa. obscuriensis* Parker 1996:4 nom. nud. 
    = Echinocereus scheeri var. obscuriensis.
Echinocereus scheeri fa. salmianus* Parker 1996:4 nom. nud. 
  = Echinocereus salmianius.
Echinocereus schereri Frank 1990a:154-159.
TL: Mexico, Durango,near Las Minas, Rio Nazas, Scherer 123 (ZSS).
Echinocereus schlechterdalii* Britton & Rose 1922:302
error for Echinocactus schlechtendalii.
Echinocereus schlini* Britton & Rose 1922:43 pro syn.
  = Echinocereus scheeri.
Echinocereus schmollii (Weingart) Taylor 1985b:140.
= Cereus schmollii Weingart 1931:251-252.
TL: Mexico, Queretero, F. Schmoll (?).
Synonyms:
Wilcoxia schmollii (Weingart) Backeberg 1935, Blätter für Kakteenforschung. 2(11):93.
Note: Wilcoxia nerispina* Backeberg 1960:2078 nom. invalid. is usually considered a
synonym of this species.
Wilcoxia schmollii var. lanata* Cartier 1980, Succulentes 2(4):8 nom. inval.
Echinocereus schmollii fa. nerispina* Parker 1991:6 nom. nud. 
  = Wilcoxia nerispina*
Echinocereus schwarzii* Backeberg 1960:2032 pro syn.
  = Echinocereus radians.
Echinocereus schwarzii Lau 1982:27-29.
TL: Mexico, Durango, near Guenacevi, 1800-2100m, Lau 1305
(POM).
Synonyms:
Echinocereus adustus var. schwarzii (Lau) Taylor 1985a:268.
Echinocereus adustus ssp. schwarzii (Lau) Taylor 1992:8
Echinocereus sciurus (K. Brandegee) K. Brandegee ex Dams 1904:130.
= Cereus sciurus K. Brandegee 1904:192.
TL: Mexico, Baja California Sur, hills near San Jose del Cabo,
    T.S. Brandegee, April 1897 (UC).
Echinocereus sciurus var. floresii (Backeberg) Taylor 1985b:115
= Echinocereus floresii.
Echinocereus sciurus ssp. floresii (Backeberg) Taylor 1997:10
= Echinocereus floresii.
Echinocereus sciurus var. floresii fa. townsendianus* Parker 1996:5 nom. nud.
= Echinocereus townsendianus*

Echinocereus scopulorum Britton & Rose 1922:30-31.
TL: Mexico, Sonora, near Guaymas, Rose et al 12570, 10 March
1910 (US).
Synonym:
Echinocereus pectinatus var. scopulorum (Britton & Rose) Marshall 1955:61.
Echinocereus scopulorum ssp. pseudopectinatus Blum & Langein Blum et al. 1998:342.
= Echinocereus pseudopectinatus.
*Echinocereus serpentinus Lemaire= Nyctocereus serpentinus.
Echinocereus setosus* Hort. ex Taylor 1988:76 pro syn.
  = Echinocereus freudenbergeri.
Cereus similis* Regel & Klein 1860:29. according to Hemsley 1880:546, but not found on that page.
Perhaps  this is the same as Cereus similis Meinshausen 1858 in Wochenschrift für Gärtnerei und
Pflanzenkunde
1:29.
TL: Mexico, Karwinsky (?)
*Echinocereus spachianus Rümpler= Echinopsis spachiana.
*Echinocereus spinibarbis Salm Dyck= Eulychnia spinibarbis.
Echinocereus spinibarbis* Haage 1900:161 nom. nud.
Echinocereus spinibarbis* Hildmann ex Fobe 1911:58 nom. illegit.
  (later homonym).
Echinocereus spinigemmatus Lau 1984:248-250.
TL: Mexico, Jalisco, near San Andres Cohamiata, 1600m, Lau
1246, 25 Apr. 1974 (ZSS AA18- 48).
Echinocereus spinosissimus Walton 1899a:162.
TL: El Paso, Texas.
Note: Usually considered a synonym of Echinocereus dasyacanthus.
Echinocereus spinosus* Coulter ex Schelle 1926:174 pro syn.
  = Echinocereus pectinatus var. armatus.
*Echinocereus splendens Lemaire
  = Nyctocereus splendens.
Echinocereus standleyi Britton & Rose 1922:24.
TL: New Mexico, Sacramento Mtns.,100 miles north of El Paso,
S. L. Pattison (US).
Synonyms:
Echinocereus viridiflorus var. standleyi (Britton & Rose)Orcutt ex Weniger 1969:37.
Echinocereus viridiflorus fa. standleyi* Parker 1996:3 nom. nud.
Echinocereus steereae Clover 1938:565-567.
TL: Texas, Chisos Mtns., Mrs Lois Steere (MICH).
Synonyms:
Echinocereus dasyacanthus var. steereae (Clover) Marshall 1945:115.
Echinocereus pectinatus var. steereae* Rümpler ex Weniger 1970:32 pro syn.
Echinocereus dasyacanthus fa. steereae* Parker 1996:2 nom. nud.
Echinocereus stoloniferus Marshall 1938:159-160.
TL: Mexico, Sonora, Mountain facing Rancho Guirocoba, 5000ft,
  Bristol & Marshall, 19 Mar. 1935 (DS 251176).
Synonyms:
Echinocereus subterraneus* Hort. ex Bailey 1941:271. pro. syn.
Echinocereus stoloniferus var. tayopensis (Marshall) Taylor 1985a:258
= Echinocereus tayopensis.
Echinocereus stoloniferus ssp. tayopensis* (Marshall) Pichler 1994:103-106. nom. inval.
  = Echinocereus tayopensis.
Echinocereus stoloniferus ssp. tayopensis (Marshall)Taylor 1997:10
= Echinocereus tayopensis.
Echinocereus stramineus (Engelmann) Engelmann ex Seitz 1870:11.
= Cereus stramineus Engelmann 1856:282.
[fuller description Engelmann 1859:35, t. 46]
TL: Texas, Mtns near El Paso, Wright, 1851 (MO).
Synonyms:
Echinocereus enneacanthus var. stramineus (Engelmann) Benson 1969d:127.
Note:Usually cited as Echinocereus stramineus Engelmann ex Rümpler 1885:797.
Echinocereus stramineus var. conglomeratus (Blanc) Bravo 1974:47
  = Echinocereus conglomeratus.
Echinocereus stramineus fa. conglomeratus* Parker 1996:3 nom. nud. 
  = Echinocereus conglomeratus.
Echinocereus stramineus var. major* Haage 1897:13 nom. nud
Echinocereus stramineus var. major* Schelle 1907:136 nom. nud.
Echinocereus stramineus var. major-aureispina* Fric 1924:121 nom. nud.
Echinocereus stramineus var. occidentalis Taylor 1988:70-71.
TL: Mexico, Durango, southeast of La Cuesta, 140 km north of city
of Durango, on volcanic rocks, 1400-1450 m., N.P. Taylor 240b,
1 July 1986 (MEXU).
Synonyms:
Echinocereus stramineus ssp. occidentalis (Taylor) Taylor
1997:10
Echinocereus stramineus ssp. occidentalis (Taylor) Taylor 1997:10
= Echinocereus stramineus var. occidentalis.
Echinocereus stramineus var. parkeri* Seed list 1996 – British Cactus
    and Succulent Journal. = Echinocereus parkeri.
Echinocereus stramineus var. ruberrimus* Fric 1924:121 nom. nud.
Echinocereus stramineus var. spinarosa* Fric 1924:121 nom. nud.
Echinocereus strausianus Haage ex Quehl 1900:70-71.
TL: Texas.
Note: Usually considered a synonym of Echinocereus viridiflorus var. cylindricus.
*Echinocereus strigosus Lemaire = Echinopsis strigosa.
Echinocereus subinermis Salm-Dyck ex Scheer 1856:291.
TL: Mexico, Chihuahua, Potts (?).
Synonyms:
Cereus subinermis (Salm-Dyck ex Scheer) Hemsley 1880:546.
Echinocereus subinermis var. aculeatus Unger 1984:164.
TL: Mexico, Chihuahua, Rio Batopilasnear la Bufa, 700m, Unger, 
1980 (ZSS AA50- 70).
Synonyms:
Echinocereus subinermis fa. aculeatus* Parker 1996:4 nom. nud.
Echinocereus subinermis fa. aculeatus* Parker 1996:4 nom. nud. 
  = Echinocereus subinermis var. aculeatus.
Echinocereus subinermis var. luteus (Britton & Rose) Knuth in
Backeberg & Knuth 1935:312
= Echinocereus luteus.
Echinocereus subinermis fa. luteus (Britton & Rose) Krainz 1967:51
= Echinocereus luteus.
Echinocereus subinermis var. ochoterenae Unger 1984:164
  = Echinocereus ochoterenae.
Echinocereus subinermis ssp. ochoterenae (Ortega) Taylor 1997:10
    = Echinocereus ochoterenae.
Echinocereus subterraneus* Hort. ex Bailey 1941:271 pro syn.
  = Echinocereus stoloniferus.
Echinocereus subterraneus* Backeberg 1960:2012 nom. inval. 
(no type designated).
TL: Mexico[See Echinocereus sanpedroensis].
Echinocereus tamaulipensis* Fric 1926:441 nom. nud.
[probably = Echinocereus pectinatus var. tamaulipensis].
Echinocereus tamaulipensis (Werdermann) Lange 1995:139- 140.
= Wilcoxia tamaulipensis Werdermann 1938:86.
TL: Mexico, Southern Coahuila, Hummel s.n. (B).
Synonyms:
Echinocereus poselgeri fa. tamaulipensis* Parker 1996:6  nom. nud.
Echinocereus tamaulipensis ssp. deherdtii Lange 1995:140–141.
TL: Mexico, Coahuila, El Sol, JE (Herbarium Hauskneckt Jena).
Synonyms:
Echinocereus poselgeri var. waldeisii fa. deherdtii* Parker 1996:6 nom. nud.
Echinocereus tamaulipensis ssp. waldeisii (Haugg) Lange 1995:142
= Echinocereus waldeisii.
Echinocereus tayopensis Marshall 1956a:78-80.
TL: Mexico, Sonora, Rancho Saucito, between Bacadehuachi &
Nacori Chico, 3500 ft (29º42’N 109º04’W), D. Gold & H.
     Sanchez-Mejorada(DES).
Synonyms:
Echinocereus stoloniferus var. tayopensis (Marshall) Taylor 1985a:258.
Echinocereus stoloniferus ssp. tayopensis (Marshall) Pichler 1994:103-106.
Echinocereus texensis Jacobi 1856:110. [not Runge 1894].
TL: Texas.
Note: This name is usually considered a synonym of Echinocereus
caespitosus.
Echinocereus texensis* Runge 1894:61-62 nom. illegit. 
(later homonym). [See Echinocereus rungei.]
Echinocereus thurberi* Monatschrift für Kakteenkunde 3:153
error for Cereus thurberi.
Echinocereus toroweapensis (Fischer) Fuersch1993:266-268
  = Echinocereus triglochidiatus var. toroweapensis.
Echinocereus townsendianus* Heyer 1989:26 nom. nud. 
  [a form of Echinocereus floresii].
Echinocereus thwaitesii* Schumann 1897:290 nom. nud.
Echinocereus trichacanthus Rümpler 1885:799.
TL: unknown
*Echinocereus trichacanthus Engelmann ex Haage 1893:23
= Cephalocereus nobilis.
Echinocereus triglochidiatus Engelmann 1848a:93.
TL: New Mexico, Wolf Creek, Wislizenus (MO).
Synonyms:
Cereus triglochidiatus (Engelmann) Engelmann 1849:50.
Echinocereus paucispinus var. triglochidiatus* Schumann1897:281 nom. illegit.
Echinocereus triglochidiatus var. acifer (Jacobi) Bravo 1978:66
= Echinocereus acifer.
Echinocereus triglochidiatus var. arizonicus (Orcutt) Benson 1969a:21
= Echinocereus arizonicus.
Echinocereus triglochidiatus var. coccineus (Engelmann) Engelmann ex 
  Marshall & Bock 1941:117
    = Echinocereus coccineus.
Echinocereus triglochidiatus var. gonacanthus (Engelmann & Bigelow)
Boissevain 1940:36
= Echinocereus gonacanthus.
Echinocereus triglochidiatus var. gurneyi Benson 1969d:126.
TL: Texas, Brewster Co., igneous rocks south of Marathon, Correll
& Benson 16,488 (POM).
Synonyms:
Echinocereus coccineus var. gurneyi (Benson) Heil & Brack1988:26.
Echinocereus coccineus ssp. gurneyi * Blum 1997 Seed List nom. nud.
Echinocereus triglochidiatus var. inermis Arp 1973:132-133
= Echinocereus coccineus var. inermis.
Echinocereus triglochidiatus var. hexaedrus (Engelmann & Bigelow) Boissevain 1940:36
= Echinocereus hexaedrus.
Echinocereus triglochidiatus var. melanacanthus (Engelmann) Benson 1944:254
    = Echinocereus coccineus var. melanacanthus.
Note: This is the name used by Benson for Echinocereus  coccineus.
Echinocereus triglochidiatus var. melanacanthus fa. inermis* Alabaster 1994:105 nom. inval. 
    = Echinocereus coccineus var. inermis.
Echinocereus triglochidiatus var. mojavensis (Engelmann & Bigelow)  Benson 1944:255
    = Echinocereus mojavensis.
Echinocereus triglochidiatus ssp. mojavensis (Engelmann & Bigelow) Blum & Lange in
Blum et al. 1998:362.
    = Echinocereus mojavensis.
Echinocereus triglochidiatus var. mojavensis fa. inermis (Schumann)  Ferguson 1989:219
= Echinocereus coccineus var. inermis.
Echinocereus triglochidiatus ssp. mojavensis fa. inermis (Schumann) Blum in Blum et al. 1998:365.
= Echinocereus coccineus var. inermis.
Echinocereus triglochidiatus var. multicostatus (Cels ex Rümpler) Marshall 1953:67
= Echinocereus multicostatus Cels ex Rümpler 1885:834.
Echinocereus triglochidiatus var. neomexicanus (Standley) Standley ex Marshall & Bock 1941:118
= Echinocereus neomexicanus.
Echinocereus triglochidiatus var. neomexicanus fa. rosei* Blum 1990:12. pro syn. 
    = Echinocereus rosei.
Echinocereus triglochidiatus var. octacanthus* Mühlenpfordt ex Marshall & Bock 1941:118
   nom.invalid.   = Echinocereus octacanthus.
Echinocereus triglochidiatus var. pacificus (Englemann ex Orcutt) Engelmann ex
   Marshall & Bock 1941:118  = Echinocereus pacificus.
Echinocereus triglochidiatus var. paucispinus (Engelmann) Engelmann ex Marshall & Bock 1941:118
    = Echinocereus paucispinus.
Echinocereus triglochidiatus var. polyacanthus (Engelmann) Benson 1944:253
    = Echinocereus polyacanthus.
Echinocereus triglochidiatus var. rosei (Wooten & Standley) Marshall 1950:63
= Echinocereus rosei.
Echinocereus triglochidiatus var. toroweapensis Fischer 1991:195.
TL: Arizona, Mohave Co.,Toroweap Point, 1340 m, P.C. Fischer 7196 (ARIZ).
Synonyms:
Echinocereus toroweapensis (Fischer) Fuersch1993:266-268.
Echinocereus triglochititus* Engelm. ex Haage 1900:162. nom. nud.
[probably an error in spelling for Echinocereus triglochidiatus].
Echinocereus trockyi* Fric 1918(8) [in advertisement] nom. nud.
Echinocereus tuberculatus* Hort. ex Haage 1900:162 nom. nud.
Echinocereus tuberosus* Poselger ex Rümpler 1885:783 nom. illegit.
  = Cereus tuberosus Poselger 1853:135 non Pfeiffer.
= Echinocereus poselgeri.
Echinocereus tulensis Bravo 1973:110-111.
TL: Mexico, Tamaulipas, 1 km. south of Tula Junctionon highway
between Huizache & Juamave, Sanchez-Mejorada 2085, 4 May
1973 (MEXU).
Synonyms:
Echinocereus cinerascens var. tulensis (Bravo) Taylor 1988:69.
Echinocereus cinerascens ssp. tulensis (Bravo) Taylor 1997:8.
Echinocereus twaitesii* Hort. ex Haage 1892:119 nom. nud.
Echinocereus uehri Blanc 1888:40.
TL: unknown
Echinocereus undulatus* Hildmann ex Schumann 1895:124 pro syn. 
  = Echinocereus cinerascens.
Echinocereus uspenskii Blanc 1888:40.
TL: unknown
Synonyms:
Echinocereus enneacanthus fa. uspenskii* Parker 1996:2 nom. nud.
Echinocereus uspenskii* Haage 1900:162 nom. illegit. (later homonym).
Echinocereus utahensis* Köhres Cat 1984. nom. nud.
Echinocereus vatteri* Botzenhart Kakteen und andere Sukkulenten
1968 front cover nom. nud. = Echinocereus viereckii.
Echinocereus viereckii Werdermann 1934:188-189.
TL: Mexico, Tamaulipas, Mtns. between Juamave & Cd. Victoria,
1500-2000m.
Echinocereus viereckii ssp. huastecensis Blum, Lange & Rutow in
Blum et al.1998:156-157.
TL: Mexico, Nuevo Leon, Rayones Canyon, 15 km west of federal
highway 85 on state road 6, Taylor 192,9June 1986 (K)
Echinocereus viereckii var. morricalii (Riha) Taylor 1985b:93
= Echinocereus morricalii.
Echinocereus viereckii ssp. morricalii (Riha) Taylor 1997:10
  = Echinocereus morricalii.
Echinocereus virickii* White 1948:288 nom. nud.
Echinocereus viridiflorus Engelmann 1848a:91.
TL: New Mexico,Wolf Creek, Wislizenus 514, (MO 2016902).
Synonyms:
Cereus viridiflorus (Engelmann) Engelmann 1849:50.
Echinocereus labouretii* Förster ex Rümpler 1885:811 pro syn.
Echinocereus viridiflorus Engelm. var. canus A.M.Powell & Weedin (2004),
Cacti Trans-Pecos 273 (-276; plates 171-172, map 36).
TL: Presidio Co., Solitario, S-facing crest of a novaculite ridge, c. 4847ft,  A.M.Powell & S.A.Powell 6012 (SRSC)
Echinocereus viridiflorus var. centrispinus* Bailey 1941:271 nom.  nud.
Echinocereus viridiflorus var. chloranthus (Engelmann) Backeberg 1960:2015
= Echinocereus chloranthus.
Echinocereus viridiflorus fa. chloranthus (Engelmann) Krainz 1967:51
= Echinocereus chloranthus.
Echinocereus viridiflorus ssp. chloranthus (Engelmann) Taylor  1997:10
= Echinocereus chloranthus.
Echinocereus viridiflorus var. chrysacanthus*  Schelle1907:129 nom. nud.
Echinocereus viridiflorus var. correllii Benson 1969d:128.
TL: Texas, Brewster Co., above Ft. Peña Colorada, 4 mi. south
of Marathon,4100 ft., Benson & Correll 16485, 4 Apr. 1965.
(POM317079).
Synonyms:
       Echinocereus viridiflorus ssp. correllii (Benson)Blum & Lange  in Blum et al. 1998:198.
Echinocereus viridiflorus ssp. correllii (Benson)Blum & Lange in  Blum et al. 1998:198.
=Echinocereus viridiflorus var. correllii
Echinocereus viridiflorus var. cylindricus (Engelmann) Engelmann
  ex Rümpler 1885:812.
= Cereus viridiflorus var. cylindricus Engelmann 1856:278.
TL: Texas, Valley of Limpia Creek, Wright, 11 June 1851
(MO 2016904).
Synonyms:
Echinocereus chloranthus var. cylindricus (Engelmann) Taylor 1984:169.
Echinocereus viridiflorus ssp. cylindricus (Engelmann) Taylor 1991:10.
Echinocereus chloranthus ssp. cylindricus  Blum & Lange  in Blum et al. 1998:205.
Note: Echinocereus strausianus is usually considered a synonym of this variety.
Echinocereus viridiflorus var. davisii (Houghton) Marshall & Bock 1941:119
 = Echinocereus davisii.
Echinocereus viridiflorus fa. davisii (Houghton) Krainz 1967:51
= Echinocereus davisii.
Echinocereus viridiflorus ssp. davisii (Houghton) Taylor 1997:10
= Echinocereus davisii.
Echinocereus viridiflorus var. elongatus* Hort. ex Backeberg 1962:3847 nom. nud.
Echinocereus viridiflorus var. faciliflorus* Hildmann ex Schelle 1907:129 nom. nud.
Echinocereus viridiflorus var. gracilispinus Hort. ex Rümpler 1885:814.
TL: unknown
Echinocereus viridiflorus var. intermedius* Backeberg 1960:2015
nom. inval. (no type designated).
Note: This plant is said to be from Texas.
Echinocereus viridiflorus var. longispinus* Schelle 1907:129 nom. nud.
Echinocereus viridiflorus var. major* Maas 1906:142 nom. nud.
Echinocereus viridiflorus var. minor* Engelmann ex Weniger 1970:14  nom. nud. 
    = Cereus viridiflorus var. minor* Engelmann 1856:278 which is a new name for Echinocereus
viridiflorus
var. viridiflorus.
Echinocereus viridiflorus var. montanus* Künzler Catalog nom. nud.
Echinocereus viridiflorus var. neocapillus Leuck 1980:35-36
= Echinocereus chloranthus var. neocapillus.
[The validity of this publication is questionable].
Echinocereus viridiflorus var. ruberrimus* Fric 1924:121 nom. nud.
Echinocereus viridiflorus var. rubrus* Fric 1924:121 nom. nud.
Echinocereus viridiflorus var. russanthus (Weniger) Leuck 1980:34
= Echinocereus russanthus.
[The validity of this publication is questionable].
Echinocereus viridiflorus var. sanguineus* Rgl. ex Schelle 1907129 nom. nud.

    [perhaps the same as Echinocereus sanguineus].
Echinocereus viridiflorus var. standleyi (Britton & Rose) Orcutt ex Weniger 1969:37
    = Echinocereus standleyi.
Echinocereus viridiflorus fa. standleyi*. Parker 1996:3 nom. nud. 
  = Echinocereus standleyi.
Echinocereus viridiflorus var. tuberculosus* Orcutt 1926:4 nom. nud.
Echinocereus viridiflorus var. tubulosus* Heller 1900:8 nom. illegit.
= Cereus viridiflorus var. tubulosus* Coulter 1896:383
new name for Echinocereus viridiforus var. cylindricus.
Echinocereus viridiflorus var. weedinii Leuck 1980:37-39
[The validity of this publication is questionable].
TL: Texas, Jeff Davis Co., top of Timber Mtn. , 6400 ft., Leuck
& Weedin 193, 13 May 1978 (OKL).
Synonyms:
Echinocereus chloranthus var. livermorensis* Kolle 1978:34 nom. inval.
(not published in printed form)
Echinocereus russanthus var. weedinii* Mesa Garden Seed  Catalog 1996 nom. nud.
Echinocereus chloranthus ssp. weedinii* Höchstätter Seed    List 1997 nom. nud.
Echinocereus russanthus ssp. weedinii Leuck ex  Blum & Lange  in Blum et al. 1998:214-216.
Echinocereus viridiflorus Engelm. var. weedinii ( Leuck ex W.Blum & Mich.Lange ) A.D.Zimmerman 2004,
Cacti Trans-Pecos 270. 2004.
Echinocereus waldeisii Haugg 1994:6.
TL: Mexico, San Luis Potosi, west of Tula, Huizache, D. Waldeis,
1988(WU).
Synonyms:
Echinocereus tamaulipensis ssp. waldeisii (Haugg) Lange 1995:142.
Echinocereus poselgeri var. waldeisii* Parker 1996:6 nom. nud.
Echinocereus webeserii* Pavey 1979: n.p. nom. nud.
[Apparently a form of Echinocereus baileyi].
Echinocereus websterianus Lindsay 1947:153.
TL: Mexico, Gulf of California, San Pedro Nolasco Is.,
Lindsay & Bool 498, 25 Feb. 1947 (DS314191).
Echinocereus weinbergii Weingart 1912:83.
TL: North America [This plant has been relocated near Sombrerete, Zacatecas, Mexico]
Synonyms:
Echinocereus pulchellus var. weinbergii (Weingart) Taylor 1985a:272.
Echinocereus pulchellus ssp. weinbergii B(Weingart) Taylor  1997:9.
Echinocereus weinbergii var. albispinus* Hort. ex Schelle 1926:175 nom. nud.
*Echinocereus whipplei var. heilii
Spelling error for Echinocactus whipplei.
Echinocereus winterianus* Kunzmann 1985:79. nom. nud.
Echinocereus yavapaiensis Baker 2006:79.
  TL: 7km NNW of Crown King, Yavapai County, Arizona.
   Baker & Lisenby 13723 (ASU238488)
Echinocereus zuniensis* Eng. & Big. ex Haage & Schmidt 1918 nom.
   nud. In advertisement opposite p. 74 Monatsschrift für
    Kakteenkunde.28(7). = Echinocereus mojavensis var. zuniensis.
Las Cruces, New Mexico 1 May 2006. Permission to copy or reprint
for noncommercial purposes is granted.

Abkürzungen & Bibliographie

Herbarium Abbreviations

(ARIZ) – University of Arizona, Tucson
(B) – Dahlem Botanical Museum, Berlin
(CA) – California Academy of Sciences, San Francisco
(DES) – Desert Botanical Garden, Phoenix
(DS) – Dudley Herbarium, California Academy of Sciences, San Francisco
(JE) – Friedrich – Schiller University, Jena
(K) – Royal Botanical Gardens, Kew
(LAM) – Natural History Museum of Los Angeles
(MEXU) – Universidad Autónoma de Mexico
(MICH) – University of Michigan, Ann Arbor
(MO) – Missouri Botanical Garden, Saint Louis
(OKL) – University of Oklahoma, Norman
(POM) – Pomona College, Claremont, California
(PRC) – Charles University, Prague
(UC) – University of California, Berkeley
(UNM) – University of New Mexico, Albuquerque
(US) – United States National Herbarium, Washington, DC
(WU) – University of Vienna
(ZSS) – Städtische Sukkulenten-Sammlung, Zürich
Other Abbreviations
Auct.? – author unknown.
CSJ – Cactus & Succulent Journal (US).
fa. – forma.
HK – Horst Künzler Catalog
Hort. – Hortus (Garden name)
nom. illegit. – name that is illegitimate according to the rules of botanical nomenclature.
nom. inval. – name that is invalid according to the rules of botanical nomenclature.
nom. nud.- a name without any description.
nom. prov. – a provisional name.
pro syn. – as a synonym for.
sp. – species
ssp. – subspecies
TL: type locality
var. – varietas
(?) – location of type specimen unknown
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* indicates work not seen.
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